Pathogenic morphological signatures of perturbations in mitochondrial-related genes revealed by pooled imaging assay

Abstract Mutations in mitochondrial-related genes underlie numerous neurodegenerative diseases, yet the significance of most variants remains uncertain concerning disease phenotypes. Several thousand genes have been shown to regulate mitochondria in eukaryotic cells, but which of these genes are nec...

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Main Authors: Colin Kremitzki, Jason Waligorski, Graham Bachman, Lina Mohammed Ali, John Bramley, Maria Vakaki, Vinay Chandrasekaran, Purva Patel, Dhruv Mathur, Paul Hime, Robi Mitra, Jeff Milbrandt, William Buchser
Format: Article
Language:English
Published: Nature Portfolio 2025-08-01
Series:npj Imaging
Online Access:https://doi.org/10.1038/s44303-025-00097-9
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author Colin Kremitzki
Jason Waligorski
Graham Bachman
Lina Mohammed Ali
John Bramley
Maria Vakaki
Vinay Chandrasekaran
Purva Patel
Dhruv Mathur
Paul Hime
Robi Mitra
Jeff Milbrandt
William Buchser
author_facet Colin Kremitzki
Jason Waligorski
Graham Bachman
Lina Mohammed Ali
John Bramley
Maria Vakaki
Vinay Chandrasekaran
Purva Patel
Dhruv Mathur
Paul Hime
Robi Mitra
Jeff Milbrandt
William Buchser
author_sort Colin Kremitzki
collection DOAJ
description Abstract Mutations in mitochondrial-related genes underlie numerous neurodegenerative diseases, yet the significance of most variants remains uncertain concerning disease phenotypes. Several thousand genes have been shown to regulate mitochondria in eukaryotic cells, but which of these genes are necessary for proper mitochondrial function and dynamics? We investigated the degree of morphological disruptions in mitochondrial gene-silenced cells to understand the genetic contribution to the expected mitochondrial phenotype and to identify potentially pathogenic variants like pathogenic mutations in MFN2. We analyzed 5835 gRNAs in a high dimensional phenotypic dataset produced by the image-based pooled analysis platform Raft-Seq. Using the MFN2-mutant cell phenotype, we identified several genes, including TMEM11, TIMM8A, NDUFAF4, NDUFAF7, and NDUFS5 (NADH ubiquinone oxidoreductase-related genes), as crucial for normal mitochondrial dynamics in human U2OS cells. Additionally, we found several missense and UTR variants within the genes SLC25A19 and ATAD3A as drivers of mitochondrial aggregation. By examining multiple features instead of a single readout, this analysis was powered to detect genes which had morphological ‘signatures’ aligned with MFN2-mutant phenotypes. Reanalysis with anomaly detection revealed other critical genes, including APOOL, MCEE, NIT, PHB, and SLC16A7, which perturb mitochondrial network morphology in a manner divergent from MFN2. These studies show causal links between gene knockouts and gene-specific variants into the assembly or maintenance of mitochondrial dynamics and can hopefully lead to a better understanding of mitochondrial related diseases.
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spelling doaj-art-ef4c8e407e07429dac352313b03dd26f2025-08-20T03:05:56ZengNature Portfolionpj Imaging2948-197X2025-08-013111310.1038/s44303-025-00097-9Pathogenic morphological signatures of perturbations in mitochondrial-related genes revealed by pooled imaging assayColin Kremitzki0Jason Waligorski1Graham Bachman2Lina Mohammed Ali3John Bramley4Maria Vakaki5Vinay Chandrasekaran6Purva Patel7Dhruv Mathur8Paul Hime9Robi Mitra10Jeff Milbrandt11William Buchser12Department of Genetics, Washington University School of MedicineDepartment of Genetics, Washington University School of MedicineDepartment of Genetics, Washington University School of MedicineDepartment of Genetics, Washington University School of MedicineDepartment of Genetics, Washington University School of MedicineDepartment of Genetics, Washington University School of MedicineDepartment of Genetics, Washington University School of MedicineDepartment of Genetics, Washington University School of MedicineDepartment of Genetics, Washington University School of MedicineDepartment of Genetics, Washington University School of MedicineDepartment of Genetics, Washington University School of MedicineDepartment of Genetics, Washington University School of MedicineDepartment of Genetics, Washington University School of MedicineAbstract Mutations in mitochondrial-related genes underlie numerous neurodegenerative diseases, yet the significance of most variants remains uncertain concerning disease phenotypes. Several thousand genes have been shown to regulate mitochondria in eukaryotic cells, but which of these genes are necessary for proper mitochondrial function and dynamics? We investigated the degree of morphological disruptions in mitochondrial gene-silenced cells to understand the genetic contribution to the expected mitochondrial phenotype and to identify potentially pathogenic variants like pathogenic mutations in MFN2. We analyzed 5835 gRNAs in a high dimensional phenotypic dataset produced by the image-based pooled analysis platform Raft-Seq. Using the MFN2-mutant cell phenotype, we identified several genes, including TMEM11, TIMM8A, NDUFAF4, NDUFAF7, and NDUFS5 (NADH ubiquinone oxidoreductase-related genes), as crucial for normal mitochondrial dynamics in human U2OS cells. Additionally, we found several missense and UTR variants within the genes SLC25A19 and ATAD3A as drivers of mitochondrial aggregation. By examining multiple features instead of a single readout, this analysis was powered to detect genes which had morphological ‘signatures’ aligned with MFN2-mutant phenotypes. Reanalysis with anomaly detection revealed other critical genes, including APOOL, MCEE, NIT, PHB, and SLC16A7, which perturb mitochondrial network morphology in a manner divergent from MFN2. These studies show causal links between gene knockouts and gene-specific variants into the assembly or maintenance of mitochondrial dynamics and can hopefully lead to a better understanding of mitochondrial related diseases.https://doi.org/10.1038/s44303-025-00097-9
spellingShingle Colin Kremitzki
Jason Waligorski
Graham Bachman
Lina Mohammed Ali
John Bramley
Maria Vakaki
Vinay Chandrasekaran
Purva Patel
Dhruv Mathur
Paul Hime
Robi Mitra
Jeff Milbrandt
William Buchser
Pathogenic morphological signatures of perturbations in mitochondrial-related genes revealed by pooled imaging assay
npj Imaging
title Pathogenic morphological signatures of perturbations in mitochondrial-related genes revealed by pooled imaging assay
title_full Pathogenic morphological signatures of perturbations in mitochondrial-related genes revealed by pooled imaging assay
title_fullStr Pathogenic morphological signatures of perturbations in mitochondrial-related genes revealed by pooled imaging assay
title_full_unstemmed Pathogenic morphological signatures of perturbations in mitochondrial-related genes revealed by pooled imaging assay
title_short Pathogenic morphological signatures of perturbations in mitochondrial-related genes revealed by pooled imaging assay
title_sort pathogenic morphological signatures of perturbations in mitochondrial related genes revealed by pooled imaging assay
url https://doi.org/10.1038/s44303-025-00097-9
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