SARS-CoV-2 ORF3a drives dynamic dense body formation for optimal viral infectivity

Abstract SARS-CoV-2 hijacks multiple organelles for virion assembly, of which the mechanisms have not been fully understood. Here, we identified a SARS-CoV-2-driven membrane structure named the 3a dense body (3DB). 3DBs are unusual electron-dense and dynamic structures driven by the accessory protei...

Full description

Saved in:
Bibliographic Details
Main Authors: Stella Hartmann, Lisa Radochonski, Chengjin Ye, Luis Martinez-Sobrido, Jueqi Chen
Format: Article
Language:English
Published: Nature Portfolio 2025-05-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-025-59475-x
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1850273456944316416
author Stella Hartmann
Lisa Radochonski
Chengjin Ye
Luis Martinez-Sobrido
Jueqi Chen
author_facet Stella Hartmann
Lisa Radochonski
Chengjin Ye
Luis Martinez-Sobrido
Jueqi Chen
author_sort Stella Hartmann
collection DOAJ
description Abstract SARS-CoV-2 hijacks multiple organelles for virion assembly, of which the mechanisms have not been fully understood. Here, we identified a SARS-CoV-2-driven membrane structure named the 3a dense body (3DB). 3DBs are unusual electron-dense and dynamic structures driven by the accessory protein ORF3a via remodeling a specific subset of the trans-Golgi network (TGN) and early endosomal membrane. 3DB formation is conserved in related bat and pangolin coronaviruses but was lost during the evolution to SARS-CoV. During SARS-CoV-2 infection, 3DB recruits the viral structural proteins spike (S) and membrane (M) and undergoes dynamic fusion/fission to maintain the optimal unprocessed-to-processed ratio of S on assembled virions. Disruption of 3DB formation resulted in virions assembled with an abnormal S processing rate, leading to a dramatic reduction in viral entry efficiency. Our study uncovers the crucial role of 3DB in maintaining maximal SARS-CoV-2 infectivity and highlights its potential as a target for COVID-19 prophylactics and therapeutics.
format Article
id doaj-art-da9d2319dd7f4d208faf9b6e7bf1bb79
institution OA Journals
issn 2041-1723
language English
publishDate 2025-05-01
publisher Nature Portfolio
record_format Article
series Nature Communications
spelling doaj-art-da9d2319dd7f4d208faf9b6e7bf1bb792025-08-20T01:51:28ZengNature PortfolioNature Communications2041-17232025-05-0116112210.1038/s41467-025-59475-xSARS-CoV-2 ORF3a drives dynamic dense body formation for optimal viral infectivityStella Hartmann0Lisa Radochonski1Chengjin Ye2Luis Martinez-Sobrido3Jueqi Chen4Department of Microbiology, University of ChicagoDepartment of Microbiology, University of ChicagoTexas Biomedical Research InstituteTexas Biomedical Research InstituteDepartment of Microbiology, University of ChicagoAbstract SARS-CoV-2 hijacks multiple organelles for virion assembly, of which the mechanisms have not been fully understood. Here, we identified a SARS-CoV-2-driven membrane structure named the 3a dense body (3DB). 3DBs are unusual electron-dense and dynamic structures driven by the accessory protein ORF3a via remodeling a specific subset of the trans-Golgi network (TGN) and early endosomal membrane. 3DB formation is conserved in related bat and pangolin coronaviruses but was lost during the evolution to SARS-CoV. During SARS-CoV-2 infection, 3DB recruits the viral structural proteins spike (S) and membrane (M) and undergoes dynamic fusion/fission to maintain the optimal unprocessed-to-processed ratio of S on assembled virions. Disruption of 3DB formation resulted in virions assembled with an abnormal S processing rate, leading to a dramatic reduction in viral entry efficiency. Our study uncovers the crucial role of 3DB in maintaining maximal SARS-CoV-2 infectivity and highlights its potential as a target for COVID-19 prophylactics and therapeutics.https://doi.org/10.1038/s41467-025-59475-x
spellingShingle Stella Hartmann
Lisa Radochonski
Chengjin Ye
Luis Martinez-Sobrido
Jueqi Chen
SARS-CoV-2 ORF3a drives dynamic dense body formation for optimal viral infectivity
Nature Communications
title SARS-CoV-2 ORF3a drives dynamic dense body formation for optimal viral infectivity
title_full SARS-CoV-2 ORF3a drives dynamic dense body formation for optimal viral infectivity
title_fullStr SARS-CoV-2 ORF3a drives dynamic dense body formation for optimal viral infectivity
title_full_unstemmed SARS-CoV-2 ORF3a drives dynamic dense body formation for optimal viral infectivity
title_short SARS-CoV-2 ORF3a drives dynamic dense body formation for optimal viral infectivity
title_sort sars cov 2 orf3a drives dynamic dense body formation for optimal viral infectivity
url https://doi.org/10.1038/s41467-025-59475-x
work_keys_str_mv AT stellahartmann sarscov2orf3adrivesdynamicdensebodyformationforoptimalviralinfectivity
AT lisaradochonski sarscov2orf3adrivesdynamicdensebodyformationforoptimalviralinfectivity
AT chengjinye sarscov2orf3adrivesdynamicdensebodyformationforoptimalviralinfectivity
AT luismartinezsobrido sarscov2orf3adrivesdynamicdensebodyformationforoptimalviralinfectivity
AT jueqichen sarscov2orf3adrivesdynamicdensebodyformationforoptimalviralinfectivity