Crossing the barrier or how regulation of ovastacin controls fertilization and translates into clinical phenotypes

Summary: The zona pellucida, a glycoprotein matrix enveloping the mammalian egg, exerts essential functions during fertilization and early embryonic development. Its safeguard property regulates sperm entry and thus indirectly controls fertility. Limited proteolysis by the metalloproteinase ovastaci...

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Main Authors: Nele von Wiegen, Christian Behl, Hagen Körschgen
Format: Article
Language:English
Published: Elsevier 2025-08-01
Series:iScience
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Online Access:http://www.sciencedirect.com/science/article/pii/S2589004225012374
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author Nele von Wiegen
Christian Behl
Hagen Körschgen
author_facet Nele von Wiegen
Christian Behl
Hagen Körschgen
author_sort Nele von Wiegen
collection DOAJ
description Summary: The zona pellucida, a glycoprotein matrix enveloping the mammalian egg, exerts essential functions during fertilization and early embryonic development. Its safeguard property regulates sperm entry and thus indirectly controls fertility. Limited proteolysis by the metalloproteinase ovastacin, released from the egg during fertilization, induces hardening of the zona pellucida. This precludes sperm entry and protects the embryo until implantation. However, ovastacin leakage before fertilization causes premature hardening and infertility if activity is not inhibited. This highlights the importance of ovastacin regulation by its endogenous inhibitor, fetuin-B. Accordingly, both loss and excessive ovastacin activity are linked to infertility. Here, we review recent discoveries on how ovastacin is precisely controlled to preserve zona pellucida permeability prior to fertilization and prevent penetration afterward. Based on these molecular mechanisms, we propose explanations for clinical phenotypes of recently discovered genetic mutations in ovastacin and discuss how modulation of ovastacin activity might be employed to regulate fertilization.
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institution Kabale University
issn 2589-0042
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spelling doaj-art-7e7e52867a84406aa3cc200796ead3c22025-08-20T03:27:52ZengElsevieriScience2589-00422025-08-0128811297610.1016/j.isci.2025.112976Crossing the barrier or how regulation of ovastacin controls fertilization and translates into clinical phenotypesNele von Wiegen0Christian Behl1Hagen Körschgen2Institute of Pathobiochemistry, University Medical Center of the Johannes Gutenberg University Mainz, Duesbergweg 6, D-55128 Mainz, GermanyInstitute of Pathobiochemistry, University Medical Center of the Johannes Gutenberg University Mainz, Duesbergweg 6, D-55128 Mainz, GermanyInstitute of Pathobiochemistry, University Medical Center of the Johannes Gutenberg University Mainz, Duesbergweg 6, D-55128 Mainz, Germany; Corresponding authorSummary: The zona pellucida, a glycoprotein matrix enveloping the mammalian egg, exerts essential functions during fertilization and early embryonic development. Its safeguard property regulates sperm entry and thus indirectly controls fertility. Limited proteolysis by the metalloproteinase ovastacin, released from the egg during fertilization, induces hardening of the zona pellucida. This precludes sperm entry and protects the embryo until implantation. However, ovastacin leakage before fertilization causes premature hardening and infertility if activity is not inhibited. This highlights the importance of ovastacin regulation by its endogenous inhibitor, fetuin-B. Accordingly, both loss and excessive ovastacin activity are linked to infertility. Here, we review recent discoveries on how ovastacin is precisely controlled to preserve zona pellucida permeability prior to fertilization and prevent penetration afterward. Based on these molecular mechanisms, we propose explanations for clinical phenotypes of recently discovered genetic mutations in ovastacin and discuss how modulation of ovastacin activity might be employed to regulate fertilization.http://www.sciencedirect.com/science/article/pii/S2589004225012374BiochemistryCell biologyDevelopmental biology
spellingShingle Nele von Wiegen
Christian Behl
Hagen Körschgen
Crossing the barrier or how regulation of ovastacin controls fertilization and translates into clinical phenotypes
iScience
Biochemistry
Cell biology
Developmental biology
title Crossing the barrier or how regulation of ovastacin controls fertilization and translates into clinical phenotypes
title_full Crossing the barrier or how regulation of ovastacin controls fertilization and translates into clinical phenotypes
title_fullStr Crossing the barrier or how regulation of ovastacin controls fertilization and translates into clinical phenotypes
title_full_unstemmed Crossing the barrier or how regulation of ovastacin controls fertilization and translates into clinical phenotypes
title_short Crossing the barrier or how regulation of ovastacin controls fertilization and translates into clinical phenotypes
title_sort crossing the barrier or how regulation of ovastacin controls fertilization and translates into clinical phenotypes
topic Biochemistry
Cell biology
Developmental biology
url http://www.sciencedirect.com/science/article/pii/S2589004225012374
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AT christianbehl crossingthebarrierorhowregulationofovastacincontrolsfertilizationandtranslatesintoclinicalphenotypes
AT hagenkorschgen crossingthebarrierorhowregulationofovastacincontrolsfertilizationandtranslatesintoclinicalphenotypes