Heparin-Binding Hemagglutinin-Induced Trained Immunity in Macrophages: Implications for Antimycobacterial Defense

Tuberculosis (TB) is a major global health threat, with the current <i>Bacillus Calmette–Guérin</i> (BCG) vaccine having limited efficacy against adult pulmonary disease. Trained immunity (TI) is a form of innate immune memory that enhances antimicrobial defense. It is characterized by t...

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Main Authors: Yongqiang Li, Xiuping Jia, Jinhua Tang, Huilian Qiao, Jiani Zhou, Yueyun Ma
Format: Article
Language:English
Published: MDPI AG 2025-07-01
Series:Biomolecules
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Online Access:https://www.mdpi.com/2218-273X/15/7/959
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author Yongqiang Li
Xiuping Jia
Jinhua Tang
Huilian Qiao
Jiani Zhou
Yueyun Ma
author_facet Yongqiang Li
Xiuping Jia
Jinhua Tang
Huilian Qiao
Jiani Zhou
Yueyun Ma
author_sort Yongqiang Li
collection DOAJ
description Tuberculosis (TB) is a major global health threat, with the current <i>Bacillus Calmette–Guérin</i> (BCG) vaccine having limited efficacy against adult pulmonary disease. Trained immunity (TI) is a form of innate immune memory that enhances antimicrobial defense. It is characterized by the epigenetic and metabolic reprogramming of innate immune cells and holds promise as a promising approach to prevent TB. In this study, we investigated the capacity of heparin-binding hemagglutinin (HBHA), a methylated antigen of <i>Mycobacterium tuberculosis</i>, to induce TI in murine RAW264.7 macrophages, human-derived THP-1 macrophages, and human peripheral blood mononuclear cells (hPBMCs). HBHA-trained macrophages exhibited the enhanced expression of pro-inflammatory cytokines (IL-1β, IL-6, TNF-α) following secondary lipopolysaccharide stimulation. The epigenetic profiling indicated elevated levels of H3K4me1 and H3K4me3 histone marks at cytokine gene loci. Further, metabolic analysis revealed heightened lactate production and the increased expression of glycolytic enzymes. Functionally, HBHA-trained macrophages exhibited improved control of intracellular mycobacteria, as evidenced by a significant reduction in colony-forming units following BCG infection. These findings elucidate that HBHA induces a functional TI phenotype via coordinated epigenetic and metabolic changes, and suggest HBHA may serve as a valuable tool for studying TI and its relevance to host defense against mycobacterial infections, pending further in vivo and clinical validation.
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spelling doaj-art-5de0647fb84b4d4ba12107b427281ed42025-08-20T03:58:27ZengMDPI AGBiomolecules2218-273X2025-07-0115795910.3390/biom15070959Heparin-Binding Hemagglutinin-Induced Trained Immunity in Macrophages: Implications for Antimycobacterial DefenseYongqiang Li0Xiuping Jia1Jinhua Tang2Huilian Qiao3Jiani Zhou4Yueyun Ma5Department of Clinical Laboratory, Air Force Medical Center, Beijing 100142, ChinaCollege of Life Science, Northwest University, Xi’an 710069, ChinaDepartment of Clinical Laboratory, Air Force Medical Center, Beijing 100142, ChinaDepartment of Clinical Laboratory, Air Force Medical Center, Beijing 100142, ChinaCollege of Life Science, Northwest University, Xi’an 710069, ChinaDepartment of Clinical Laboratory, Air Force Medical Center, Beijing 100142, ChinaTuberculosis (TB) is a major global health threat, with the current <i>Bacillus Calmette–Guérin</i> (BCG) vaccine having limited efficacy against adult pulmonary disease. Trained immunity (TI) is a form of innate immune memory that enhances antimicrobial defense. It is characterized by the epigenetic and metabolic reprogramming of innate immune cells and holds promise as a promising approach to prevent TB. In this study, we investigated the capacity of heparin-binding hemagglutinin (HBHA), a methylated antigen of <i>Mycobacterium tuberculosis</i>, to induce TI in murine RAW264.7 macrophages, human-derived THP-1 macrophages, and human peripheral blood mononuclear cells (hPBMCs). HBHA-trained macrophages exhibited the enhanced expression of pro-inflammatory cytokines (IL-1β, IL-6, TNF-α) following secondary lipopolysaccharide stimulation. The epigenetic profiling indicated elevated levels of H3K4me1 and H3K4me3 histone marks at cytokine gene loci. Further, metabolic analysis revealed heightened lactate production and the increased expression of glycolytic enzymes. Functionally, HBHA-trained macrophages exhibited improved control of intracellular mycobacteria, as evidenced by a significant reduction in colony-forming units following BCG infection. These findings elucidate that HBHA induces a functional TI phenotype via coordinated epigenetic and metabolic changes, and suggest HBHA may serve as a valuable tool for studying TI and its relevance to host defense against mycobacterial infections, pending further in vivo and clinical validation.https://www.mdpi.com/2218-273X/15/7/959heparin-binding hemagglutinintrained immunitymacrophagetuberculosisepigenetic reprogram
spellingShingle Yongqiang Li
Xiuping Jia
Jinhua Tang
Huilian Qiao
Jiani Zhou
Yueyun Ma
Heparin-Binding Hemagglutinin-Induced Trained Immunity in Macrophages: Implications for Antimycobacterial Defense
Biomolecules
heparin-binding hemagglutinin
trained immunity
macrophage
tuberculosis
epigenetic reprogram
title Heparin-Binding Hemagglutinin-Induced Trained Immunity in Macrophages: Implications for Antimycobacterial Defense
title_full Heparin-Binding Hemagglutinin-Induced Trained Immunity in Macrophages: Implications for Antimycobacterial Defense
title_fullStr Heparin-Binding Hemagglutinin-Induced Trained Immunity in Macrophages: Implications for Antimycobacterial Defense
title_full_unstemmed Heparin-Binding Hemagglutinin-Induced Trained Immunity in Macrophages: Implications for Antimycobacterial Defense
title_short Heparin-Binding Hemagglutinin-Induced Trained Immunity in Macrophages: Implications for Antimycobacterial Defense
title_sort heparin binding hemagglutinin induced trained immunity in macrophages implications for antimycobacterial defense
topic heparin-binding hemagglutinin
trained immunity
macrophage
tuberculosis
epigenetic reprogram
url https://www.mdpi.com/2218-273X/15/7/959
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