The degree of polymerization and sulfation patterns in heparan sulfate are critical determinants of cytomegalovirus entry into host cells.

Several enveloped viruses, including herpesviruses attach to host cells by initially interacting with cell surface heparan sulfate (HS) proteoglycans followed by specific coreceptor engagement which culminates in virus-host membrane fusion and virus entry. Interfering with HS-herpesvirus interaction...

Full description

Saved in:
Bibliographic Details
Main Authors: Dipanwita Mitra, Mohammad H Hasan, John T Bates, Michael A Bierdeman, Dallas R Ederer, Rinkuben C Parmar, Lauren A Fassero, Quntao Liang, Hong Qiu, Vaibhav Tiwari, Fuming Zhang, Robert J Linhardt, Joshua S Sharp, Lianchun Wang, Ritesh Tandon
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2021-08-01
Series:PLoS Pathogens
Online Access:https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1009803&type=printable
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1850159958912401408
author Dipanwita Mitra
Mohammad H Hasan
John T Bates
Michael A Bierdeman
Dallas R Ederer
Rinkuben C Parmar
Lauren A Fassero
Quntao Liang
Hong Qiu
Vaibhav Tiwari
Fuming Zhang
Robert J Linhardt
Joshua S Sharp
Lianchun Wang
Ritesh Tandon
author_facet Dipanwita Mitra
Mohammad H Hasan
John T Bates
Michael A Bierdeman
Dallas R Ederer
Rinkuben C Parmar
Lauren A Fassero
Quntao Liang
Hong Qiu
Vaibhav Tiwari
Fuming Zhang
Robert J Linhardt
Joshua S Sharp
Lianchun Wang
Ritesh Tandon
author_sort Dipanwita Mitra
collection DOAJ
description Several enveloped viruses, including herpesviruses attach to host cells by initially interacting with cell surface heparan sulfate (HS) proteoglycans followed by specific coreceptor engagement which culminates in virus-host membrane fusion and virus entry. Interfering with HS-herpesvirus interactions has long been known to result in significant reduction in virus infectivity indicating that HS play important roles in initiating virus entry. In this study, we provide a series of evidence to prove that specific sulfations as well as the degree of polymerization (dp) of HS govern human cytomegalovirus (CMV) binding and infection. First, purified CMV extracellular virions preferentially bind to sulfated longer chain HS on a glycoarray compared to a variety of unsulfated glycosaminoglycans including unsulfated shorter chain HS. Second, the fraction of glycosaminoglycans (GAG) displaying higher dp and sulfation has a larger impact on CMV titers compared to other fractions. Third, cell lines deficient in specific glucosaminyl sulfotransferases produce significantly reduced CMV titers compared to wild-type cells and virus entry is compromised in these mutant cells. Finally, purified glycoprotein B shows strong binding to heparin, and desulfated heparin analogs compete poorly with heparin for gB binding. Taken together, these results highlight the significance of HS chain length and sulfation patterns in CMV attachment and infectivity.
format Article
id doaj-art-494ffbd93696441bb2b9787e8edc5fbf
institution OA Journals
issn 1553-7366
1553-7374
language English
publishDate 2021-08-01
publisher Public Library of Science (PLoS)
record_format Article
series PLoS Pathogens
spelling doaj-art-494ffbd93696441bb2b9787e8edc5fbf2025-08-20T02:23:18ZengPublic Library of Science (PLoS)PLoS Pathogens1553-73661553-73742021-08-01178e100980310.1371/journal.ppat.1009803The degree of polymerization and sulfation patterns in heparan sulfate are critical determinants of cytomegalovirus entry into host cells.Dipanwita MitraMohammad H HasanJohn T BatesMichael A BierdemanDallas R EdererRinkuben C ParmarLauren A FasseroQuntao LiangHong QiuVaibhav TiwariFuming ZhangRobert J LinhardtJoshua S SharpLianchun WangRitesh TandonSeveral enveloped viruses, including herpesviruses attach to host cells by initially interacting with cell surface heparan sulfate (HS) proteoglycans followed by specific coreceptor engagement which culminates in virus-host membrane fusion and virus entry. Interfering with HS-herpesvirus interactions has long been known to result in significant reduction in virus infectivity indicating that HS play important roles in initiating virus entry. In this study, we provide a series of evidence to prove that specific sulfations as well as the degree of polymerization (dp) of HS govern human cytomegalovirus (CMV) binding and infection. First, purified CMV extracellular virions preferentially bind to sulfated longer chain HS on a glycoarray compared to a variety of unsulfated glycosaminoglycans including unsulfated shorter chain HS. Second, the fraction of glycosaminoglycans (GAG) displaying higher dp and sulfation has a larger impact on CMV titers compared to other fractions. Third, cell lines deficient in specific glucosaminyl sulfotransferases produce significantly reduced CMV titers compared to wild-type cells and virus entry is compromised in these mutant cells. Finally, purified glycoprotein B shows strong binding to heparin, and desulfated heparin analogs compete poorly with heparin for gB binding. Taken together, these results highlight the significance of HS chain length and sulfation patterns in CMV attachment and infectivity.https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1009803&type=printable
spellingShingle Dipanwita Mitra
Mohammad H Hasan
John T Bates
Michael A Bierdeman
Dallas R Ederer
Rinkuben C Parmar
Lauren A Fassero
Quntao Liang
Hong Qiu
Vaibhav Tiwari
Fuming Zhang
Robert J Linhardt
Joshua S Sharp
Lianchun Wang
Ritesh Tandon
The degree of polymerization and sulfation patterns in heparan sulfate are critical determinants of cytomegalovirus entry into host cells.
PLoS Pathogens
title The degree of polymerization and sulfation patterns in heparan sulfate are critical determinants of cytomegalovirus entry into host cells.
title_full The degree of polymerization and sulfation patterns in heparan sulfate are critical determinants of cytomegalovirus entry into host cells.
title_fullStr The degree of polymerization and sulfation patterns in heparan sulfate are critical determinants of cytomegalovirus entry into host cells.
title_full_unstemmed The degree of polymerization and sulfation patterns in heparan sulfate are critical determinants of cytomegalovirus entry into host cells.
title_short The degree of polymerization and sulfation patterns in heparan sulfate are critical determinants of cytomegalovirus entry into host cells.
title_sort degree of polymerization and sulfation patterns in heparan sulfate are critical determinants of cytomegalovirus entry into host cells
url https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1009803&type=printable
work_keys_str_mv AT dipanwitamitra thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT mohammadhhasan thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT johntbates thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT michaelabierdeman thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT dallasrederer thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT rinkubencparmar thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT laurenafassero thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT quntaoliang thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT hongqiu thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT vaibhavtiwari thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT fumingzhang thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT robertjlinhardt thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT joshuassharp thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT lianchunwang thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT riteshtandon thedegreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT dipanwitamitra degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT mohammadhhasan degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT johntbates degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT michaelabierdeman degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT dallasrederer degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT rinkubencparmar degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT laurenafassero degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT quntaoliang degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT hongqiu degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT vaibhavtiwari degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT fumingzhang degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT robertjlinhardt degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT joshuassharp degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT lianchunwang degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells
AT riteshtandon degreeofpolymerizationandsulfationpatternsinheparansulfatearecriticaldeterminantsofcytomegalovirusentryintohostcells