Signaling Centers Drive Brachyury Dynamics and Lineage Commitment in mESC Aggregates

ABSTRACT Stem cell‐based models of embryogenesis have exploded in popularity, resulting in protocols with overlapping use of some reagents and differential use of others. As such, the precise contributions of individual signaling molecules, such as Chiron and BMP4, applied to whole or part of aggreg...

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Main Authors: A. Rabeling, M. Goolam
Format: Article
Language:English
Published: Wiley 2025-06-01
Series:FASEB BioAdvances
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Online Access:https://doi.org/10.1096/fba.2024-00216
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author A. Rabeling
M. Goolam
author_facet A. Rabeling
M. Goolam
author_sort A. Rabeling
collection DOAJ
description ABSTRACT Stem cell‐based models of embryogenesis have exploded in popularity, resulting in protocols with overlapping use of some reagents and differential use of others. As such, the precise contributions of individual signaling molecules, such as Chiron and BMP4, applied to whole or part of aggregates, and matrices, such as Matrigel, to the development of these models are unknown. Furthermore, the use of these different methods, signaling molecules, and matrices has yet to be directly compared under the same conditions. In this paper, we used a mouse embryonic stem cell aggregate model to compare the use of Chiron and BMP4 signaling as pulses (applied to the whole aggregate) or signaling centers (applied to part of the aggregate) and embedded them in low‐percentage Matrigel. Each factor had different effects on morphology, Brachyury protein expression, and lineage commitment, with signaling centers having different effects to pulses. BMP4 as a pulse was shown to drive neural differentiation, while signaling centers resulted in better recapitulation of aspects of anterior–posterior axis formation, with polarization of Brachyury protein and expression of anterior and posterior genes observed. This further elucidates the contributions of Chiron and BMP4 to aggregate development to better inform decisions around experimental conditions for in vitro models of embryonic development.
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spelling doaj-art-486da0d2a99841549e48c8dd20134a202025-08-20T03:46:04ZengWileyFASEB BioAdvances2573-98322025-06-0176n/an/a10.1096/fba.2024-00216Signaling Centers Drive Brachyury Dynamics and Lineage Commitment in mESC AggregatesA. Rabeling0M. Goolam1Department of Human Biology, Faculty of Health Sciences University of Cape Town Cape Town South AfricaDepartment of Human Biology, Faculty of Health Sciences University of Cape Town Cape Town South AfricaABSTRACT Stem cell‐based models of embryogenesis have exploded in popularity, resulting in protocols with overlapping use of some reagents and differential use of others. As such, the precise contributions of individual signaling molecules, such as Chiron and BMP4, applied to whole or part of aggregates, and matrices, such as Matrigel, to the development of these models are unknown. Furthermore, the use of these different methods, signaling molecules, and matrices has yet to be directly compared under the same conditions. In this paper, we used a mouse embryonic stem cell aggregate model to compare the use of Chiron and BMP4 signaling as pulses (applied to the whole aggregate) or signaling centers (applied to part of the aggregate) and embedded them in low‐percentage Matrigel. Each factor had different effects on morphology, Brachyury protein expression, and lineage commitment, with signaling centers having different effects to pulses. BMP4 as a pulse was shown to drive neural differentiation, while signaling centers resulted in better recapitulation of aspects of anterior–posterior axis formation, with polarization of Brachyury protein and expression of anterior and posterior genes observed. This further elucidates the contributions of Chiron and BMP4 to aggregate development to better inform decisions around experimental conditions for in vitro models of embryonic development.https://doi.org/10.1096/fba.2024-00216bone morphogenic protein 4brachyury proteinembryonic and fetal developmentextracellular matrixgastrulationin situ hybridization fluorescence
spellingShingle A. Rabeling
M. Goolam
Signaling Centers Drive Brachyury Dynamics and Lineage Commitment in mESC Aggregates
FASEB BioAdvances
bone morphogenic protein 4
brachyury protein
embryonic and fetal development
extracellular matrix
gastrulation
in situ hybridization fluorescence
title Signaling Centers Drive Brachyury Dynamics and Lineage Commitment in mESC Aggregates
title_full Signaling Centers Drive Brachyury Dynamics and Lineage Commitment in mESC Aggregates
title_fullStr Signaling Centers Drive Brachyury Dynamics and Lineage Commitment in mESC Aggregates
title_full_unstemmed Signaling Centers Drive Brachyury Dynamics and Lineage Commitment in mESC Aggregates
title_short Signaling Centers Drive Brachyury Dynamics and Lineage Commitment in mESC Aggregates
title_sort signaling centers drive brachyury dynamics and lineage commitment in mesc aggregates
topic bone morphogenic protein 4
brachyury protein
embryonic and fetal development
extracellular matrix
gastrulation
in situ hybridization fluorescence
url https://doi.org/10.1096/fba.2024-00216
work_keys_str_mv AT arabeling signalingcentersdrivebrachyurydynamicsandlineagecommitmentinmescaggregates
AT mgoolam signalingcentersdrivebrachyurydynamicsandlineagecommitmentinmescaggregates