Refining fine-mapping: Effect sizes and regional heritability.

Recent statistical approaches have shown that the set of all available genetic variants explains considerably more phenotypic variance of complex traits and diseases than the individual variants that are robustly associated with these phenotypes. However, rapidly increasing sample sizes constantly i...

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Main Authors: Christian Benner, Anubha Mahajan, Matti Pirinen
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2025-01-01
Series:PLoS Genetics
Online Access:https://doi.org/10.1371/journal.pgen.1011480
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author Christian Benner
Anubha Mahajan
Matti Pirinen
author_facet Christian Benner
Anubha Mahajan
Matti Pirinen
author_sort Christian Benner
collection DOAJ
description Recent statistical approaches have shown that the set of all available genetic variants explains considerably more phenotypic variance of complex traits and diseases than the individual variants that are robustly associated with these phenotypes. However, rapidly increasing sample sizes constantly improve detection and prioritization of individual variants driving the associations between genomic regions and phenotypes. Therefore, it is useful to routinely estimate how much phenotypic variance the detected variants explain for each region by taking into account the correlation structure of variants and the uncertainty in their causal status. Here we extend the FINEMAP software to estimate the effect sizes and regional heritability under the probabilistic model that assumes a handful of causal variants per region. Using the UK Biobank (UKB) data to simulate genomic regions, we demonstrate that FINEMAP provides higher precision and enables more detailed decomposition of regional heritability into individual variants than the variance component model implemented in BOLT or the fixed-effect model implemented in HESS, particularly when there are only a few causal variants in the fine-mapped region. Using data from 2,940 plasma proteins from the UKB study, we observed that on average FINEMAP identified 2.5 causal variants at an association signal and captured 36% more regional heritability than the variant with the lowest P-value. We estimate that in genomic regions with notable contribution to the total heritability, FINEMAP captures on average 13% and 40% more heritability than BOLT and HESS respectively. Our analysis shows how FINEMAP, BOLT and HESS relate to each other in cases where inference of a variant-level picture of the regional genetic architecture is possible.
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spelling doaj-art-32ccdb2b14ea41a78d4917f68b4547712025-08-20T02:28:07ZengPublic Library of Science (PLoS)PLoS Genetics1553-73901553-74042025-01-01211e101148010.1371/journal.pgen.1011480Refining fine-mapping: Effect sizes and regional heritability.Christian BennerAnubha MahajanMatti PirinenRecent statistical approaches have shown that the set of all available genetic variants explains considerably more phenotypic variance of complex traits and diseases than the individual variants that are robustly associated with these phenotypes. However, rapidly increasing sample sizes constantly improve detection and prioritization of individual variants driving the associations between genomic regions and phenotypes. Therefore, it is useful to routinely estimate how much phenotypic variance the detected variants explain for each region by taking into account the correlation structure of variants and the uncertainty in their causal status. Here we extend the FINEMAP software to estimate the effect sizes and regional heritability under the probabilistic model that assumes a handful of causal variants per region. Using the UK Biobank (UKB) data to simulate genomic regions, we demonstrate that FINEMAP provides higher precision and enables more detailed decomposition of regional heritability into individual variants than the variance component model implemented in BOLT or the fixed-effect model implemented in HESS, particularly when there are only a few causal variants in the fine-mapped region. Using data from 2,940 plasma proteins from the UKB study, we observed that on average FINEMAP identified 2.5 causal variants at an association signal and captured 36% more regional heritability than the variant with the lowest P-value. We estimate that in genomic regions with notable contribution to the total heritability, FINEMAP captures on average 13% and 40% more heritability than BOLT and HESS respectively. Our analysis shows how FINEMAP, BOLT and HESS relate to each other in cases where inference of a variant-level picture of the regional genetic architecture is possible.https://doi.org/10.1371/journal.pgen.1011480
spellingShingle Christian Benner
Anubha Mahajan
Matti Pirinen
Refining fine-mapping: Effect sizes and regional heritability.
PLoS Genetics
title Refining fine-mapping: Effect sizes and regional heritability.
title_full Refining fine-mapping: Effect sizes and regional heritability.
title_fullStr Refining fine-mapping: Effect sizes and regional heritability.
title_full_unstemmed Refining fine-mapping: Effect sizes and regional heritability.
title_short Refining fine-mapping: Effect sizes and regional heritability.
title_sort refining fine mapping effect sizes and regional heritability
url https://doi.org/10.1371/journal.pgen.1011480
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