Improving the phenotype predictions of a yeast genome‐scale metabolic model by incorporating enzymatic constraints

Abstract Genome‐scale metabolic models (GEMs) are widely used to calculate metabolic phenotypes. They rely on defining a set of constraints, the most common of which is that the production of metabolites and/or growth are limited by the carbon source uptake rate. However, enzyme abundances and kinet...

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Main Authors: Benjamín J Sánchez, Cheng Zhang, Avlant Nilsson, Petri‐Jaan Lahtvee, Eduard J Kerkhoven, Jens Nielsen
Format: Article
Language:English
Published: Springer Nature 2017-08-01
Series:Molecular Systems Biology
Subjects:
Online Access:https://doi.org/10.15252/msb.20167411
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author Benjamín J Sánchez
Cheng Zhang
Avlant Nilsson
Petri‐Jaan Lahtvee
Eduard J Kerkhoven
Jens Nielsen
author_facet Benjamín J Sánchez
Cheng Zhang
Avlant Nilsson
Petri‐Jaan Lahtvee
Eduard J Kerkhoven
Jens Nielsen
author_sort Benjamín J Sánchez
collection DOAJ
description Abstract Genome‐scale metabolic models (GEMs) are widely used to calculate metabolic phenotypes. They rely on defining a set of constraints, the most common of which is that the production of metabolites and/or growth are limited by the carbon source uptake rate. However, enzyme abundances and kinetics, which act as limitations on metabolic fluxes, are not taken into account. Here, we present GECKO, a method that enhances a GEM to account for enzymes as part of reactions, thereby ensuring that each metabolic flux does not exceed its maximum capacity, equal to the product of the enzyme's abundance and turnover number. We applied GECKO to a Saccharomyces cerevisiae GEM and demonstrated that the new model could correctly describe phenotypes that the previous model could not, particularly under high enzymatic pressure conditions, such as yeast growing on different carbon sources in excess, coping with stress, or overexpressing a specific pathway. GECKO also allows to directly integrate quantitative proteomics data; by doing so, we significantly reduced flux variability of the model, in over 60% of metabolic reactions. Additionally, the model gives insight into the distribution of enzyme usage between and within metabolic pathways. The developed method and model are expected to increase the use of model‐based design in metabolic engineering.
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spelling doaj-art-2104f48007de4f02904b64ab55859a4b2025-08-20T03:06:10ZengSpringer NatureMolecular Systems Biology1744-42922017-08-0113811610.15252/msb.20167411Improving the phenotype predictions of a yeast genome‐scale metabolic model by incorporating enzymatic constraintsBenjamín J Sánchez0Cheng Zhang1Avlant Nilsson2Petri‐Jaan Lahtvee3Eduard J Kerkhoven4Jens Nielsen5Department of Biology and Biological Engineering, Chalmers University of TechnologyScience for Life Laboratory, KTH – Royal Institute of TechnologyDepartment of Biology and Biological Engineering, Chalmers University of TechnologyDepartment of Biology and Biological Engineering, Chalmers University of TechnologyDepartment of Biology and Biological Engineering, Chalmers University of TechnologyDepartment of Biology and Biological Engineering, Chalmers University of TechnologyAbstract Genome‐scale metabolic models (GEMs) are widely used to calculate metabolic phenotypes. They rely on defining a set of constraints, the most common of which is that the production of metabolites and/or growth are limited by the carbon source uptake rate. However, enzyme abundances and kinetics, which act as limitations on metabolic fluxes, are not taken into account. Here, we present GECKO, a method that enhances a GEM to account for enzymes as part of reactions, thereby ensuring that each metabolic flux does not exceed its maximum capacity, equal to the product of the enzyme's abundance and turnover number. We applied GECKO to a Saccharomyces cerevisiae GEM and demonstrated that the new model could correctly describe phenotypes that the previous model could not, particularly under high enzymatic pressure conditions, such as yeast growing on different carbon sources in excess, coping with stress, or overexpressing a specific pathway. GECKO also allows to directly integrate quantitative proteomics data; by doing so, we significantly reduced flux variability of the model, in over 60% of metabolic reactions. Additionally, the model gives insight into the distribution of enzyme usage between and within metabolic pathways. The developed method and model are expected to increase the use of model‐based design in metabolic engineering.https://doi.org/10.15252/msb.20167411enzyme kineticsflux balance analysismolecular crowdingproteomicsSaccharomyces cerevisiae
spellingShingle Benjamín J Sánchez
Cheng Zhang
Avlant Nilsson
Petri‐Jaan Lahtvee
Eduard J Kerkhoven
Jens Nielsen
Improving the phenotype predictions of a yeast genome‐scale metabolic model by incorporating enzymatic constraints
Molecular Systems Biology
enzyme kinetics
flux balance analysis
molecular crowding
proteomics
Saccharomyces cerevisiae
title Improving the phenotype predictions of a yeast genome‐scale metabolic model by incorporating enzymatic constraints
title_full Improving the phenotype predictions of a yeast genome‐scale metabolic model by incorporating enzymatic constraints
title_fullStr Improving the phenotype predictions of a yeast genome‐scale metabolic model by incorporating enzymatic constraints
title_full_unstemmed Improving the phenotype predictions of a yeast genome‐scale metabolic model by incorporating enzymatic constraints
title_short Improving the phenotype predictions of a yeast genome‐scale metabolic model by incorporating enzymatic constraints
title_sort improving the phenotype predictions of a yeast genome scale metabolic model by incorporating enzymatic constraints
topic enzyme kinetics
flux balance analysis
molecular crowding
proteomics
Saccharomyces cerevisiae
url https://doi.org/10.15252/msb.20167411
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AT avlantnilsson improvingthephenotypepredictionsofayeastgenomescalemetabolicmodelbyincorporatingenzymaticconstraints
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