Foxn1 Is Dynamically Regulated in Thymic Epithelial Cells during Embryogenesis and at the Onset of Thymic Involution.

Thymus function requires extensive cross-talk between developing T-cells and the thymic epithelium, which consists of cortical and medullary TEC. The transcription factor FOXN1 is the master regulator of TEC differentiation and function, and declining Foxn1 expression with age results in stereotypic...

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Main Authors: Kathy E O'Neill, Nicholas Bredenkamp, Christin Tischner, Harsh J Vaidya, Frances H Stenhouse, C Diana Peddie, Craig S Nowell, Terri Gaskell, C Clare Blackburn
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2016-01-01
Series:PLoS ONE
Online Access:https://doi.org/10.1371/journal.pone.0151666
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author Kathy E O'Neill
Nicholas Bredenkamp
Christin Tischner
Harsh J Vaidya
Frances H Stenhouse
C Diana Peddie
Craig S Nowell
Terri Gaskell
C Clare Blackburn
author_facet Kathy E O'Neill
Nicholas Bredenkamp
Christin Tischner
Harsh J Vaidya
Frances H Stenhouse
C Diana Peddie
Craig S Nowell
Terri Gaskell
C Clare Blackburn
author_sort Kathy E O'Neill
collection DOAJ
description Thymus function requires extensive cross-talk between developing T-cells and the thymic epithelium, which consists of cortical and medullary TEC. The transcription factor FOXN1 is the master regulator of TEC differentiation and function, and declining Foxn1 expression with age results in stereotypical thymic involution. Understanding of the dynamics of Foxn1 expression is, however, limited by a lack of single cell resolution data. We have generated a novel reporter of Foxn1 expression, Foxn1G, to monitor changes in Foxn1 expression during embryogenesis and involution. Our data reveal that early differentiation and maturation of cortical and medullary TEC coincides with precise sub-lineage-specific regulation of Foxn1 expression levels. We further show that initiation of thymic involution is associated with reduced cTEC functionality, and proportional expansion of FOXN1-negative TEC in both cortical and medullary sub-lineages. Cortex-specific down-regulation of Foxn1 between 1 and 3 months of age may therefore be a key driver of the early stages of age-related thymic involution.
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language English
publishDate 2016-01-01
publisher Public Library of Science (PLoS)
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spelling doaj-art-16c761d49bd44448a38f17c9ffa3a3582025-08-20T02:34:12ZengPublic Library of Science (PLoS)PLoS ONE1932-62032016-01-01113e015166610.1371/journal.pone.0151666Foxn1 Is Dynamically Regulated in Thymic Epithelial Cells during Embryogenesis and at the Onset of Thymic Involution.Kathy E O'NeillNicholas BredenkampChristin TischnerHarsh J VaidyaFrances H StenhouseC Diana PeddieCraig S NowellTerri GaskellC Clare BlackburnThymus function requires extensive cross-talk between developing T-cells and the thymic epithelium, which consists of cortical and medullary TEC. The transcription factor FOXN1 is the master regulator of TEC differentiation and function, and declining Foxn1 expression with age results in stereotypical thymic involution. Understanding of the dynamics of Foxn1 expression is, however, limited by a lack of single cell resolution data. We have generated a novel reporter of Foxn1 expression, Foxn1G, to monitor changes in Foxn1 expression during embryogenesis and involution. Our data reveal that early differentiation and maturation of cortical and medullary TEC coincides with precise sub-lineage-specific regulation of Foxn1 expression levels. We further show that initiation of thymic involution is associated with reduced cTEC functionality, and proportional expansion of FOXN1-negative TEC in both cortical and medullary sub-lineages. Cortex-specific down-regulation of Foxn1 between 1 and 3 months of age may therefore be a key driver of the early stages of age-related thymic involution.https://doi.org/10.1371/journal.pone.0151666
spellingShingle Kathy E O'Neill
Nicholas Bredenkamp
Christin Tischner
Harsh J Vaidya
Frances H Stenhouse
C Diana Peddie
Craig S Nowell
Terri Gaskell
C Clare Blackburn
Foxn1 Is Dynamically Regulated in Thymic Epithelial Cells during Embryogenesis and at the Onset of Thymic Involution.
PLoS ONE
title Foxn1 Is Dynamically Regulated in Thymic Epithelial Cells during Embryogenesis and at the Onset of Thymic Involution.
title_full Foxn1 Is Dynamically Regulated in Thymic Epithelial Cells during Embryogenesis and at the Onset of Thymic Involution.
title_fullStr Foxn1 Is Dynamically Regulated in Thymic Epithelial Cells during Embryogenesis and at the Onset of Thymic Involution.
title_full_unstemmed Foxn1 Is Dynamically Regulated in Thymic Epithelial Cells during Embryogenesis and at the Onset of Thymic Involution.
title_short Foxn1 Is Dynamically Regulated in Thymic Epithelial Cells during Embryogenesis and at the Onset of Thymic Involution.
title_sort foxn1 is dynamically regulated in thymic epithelial cells during embryogenesis and at the onset of thymic involution
url https://doi.org/10.1371/journal.pone.0151666
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