Exploring the impact of forage-to-concentrate ratios on the ruminal bacteriome in vitro focusing on ciliate-associated bacteria

Abstract Ruminal ciliates are linked to methane production and nitrogen utilization efficiency in ruminants due to their association with other ruminal microorganisms. However, research on the specific interplay between ruminal bacteria and ciliates is still limited, particularly in different dietar...

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Main Authors: Geonwoo Kim, Woohyung Lee, Tansol Park
Format: Article
Language:English
Published: Nature Portfolio 2025-07-01
Series:Scientific Reports
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Online Access:https://doi.org/10.1038/s41598-025-10599-6
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author Geonwoo Kim
Woohyung Lee
Tansol Park
author_facet Geonwoo Kim
Woohyung Lee
Tansol Park
author_sort Geonwoo Kim
collection DOAJ
description Abstract Ruminal ciliates are linked to methane production and nitrogen utilization efficiency in ruminants due to their association with other ruminal microorganisms. However, research on the specific interplay between ruminal bacteria and ciliates is still limited, particularly in different dietary conditions. This study examines the effect of the forage-to-concentrate (F:C) ratio on the ruminal bacteriome in vitro, focusing on bacteria associated with Isotricha spp. and small entodinia. The rumen fluid used as the inoculum for this experiment was collected from two cannulated Hanwoo cows. Dietary treatments included high-forage (HF, F:C of 7:3), high-concentrate (HC, F:C of 3:7), and control (CON, F:C of 5:5). After 24-hour incubation, fractions for entodinia-associated bacteria (EAB), Isotricha-associated bacteria (IAB), and total bacteria (TB) were collected for bacteriome analysis using QIIME2 with full-length 16S rRNA gene sequences on the PacBio system. All fermentation parameters, except for NH3-N, showed linear changes with increasing F:C ratios (p ≤ 0.05). F:C ratio affected Isotricha spp. and Dasytricha spp. counts. Ciliate-associated bacterial fractions were significantly less diverse than the total bacterial group, as indicated by richness, phylogenetic diversity, and evenness indices. This suggests potential specific associations within ciliate-provided microhabitats. Both diet and ciliate fractions significantly influenced the overall bacteriome (p ≤ 0.05). More bacteriome features were differentially abundant due to the ciliate fraction effect rather than diet (q ≤ 0.05). Our newly proposed washing procedure, using higher ciliate cell counts and minimal bacterial contamination, effectively removed free-living or loosely associated bacteria. This allows focus on ciliate-associated bacterial populations, which may include potential symbionts or engulfed bacteria of host ruminal ciliates. Verifying these associations could provide insights into rumen microbiome dynamics, nitrogen utilization, hydrogen balance, and microbiome variation under different F:C ratios.
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spelling doaj-art-10e46e0afbb241219fce8a374b9c39bf2025-08-20T03:46:05ZengNature PortfolioScientific Reports2045-23222025-07-0115111710.1038/s41598-025-10599-6Exploring the impact of forage-to-concentrate ratios on the ruminal bacteriome in vitro focusing on ciliate-associated bacteriaGeonwoo Kim0Woohyung Lee1Tansol Park2Department of Animal Science and Technology, Chung-Ang UniversityDepartment of Animal Science and Technology, Chung-Ang UniversityDepartment of Animal Science and Technology, Chung-Ang UniversityAbstract Ruminal ciliates are linked to methane production and nitrogen utilization efficiency in ruminants due to their association with other ruminal microorganisms. However, research on the specific interplay between ruminal bacteria and ciliates is still limited, particularly in different dietary conditions. This study examines the effect of the forage-to-concentrate (F:C) ratio on the ruminal bacteriome in vitro, focusing on bacteria associated with Isotricha spp. and small entodinia. The rumen fluid used as the inoculum for this experiment was collected from two cannulated Hanwoo cows. Dietary treatments included high-forage (HF, F:C of 7:3), high-concentrate (HC, F:C of 3:7), and control (CON, F:C of 5:5). After 24-hour incubation, fractions for entodinia-associated bacteria (EAB), Isotricha-associated bacteria (IAB), and total bacteria (TB) were collected for bacteriome analysis using QIIME2 with full-length 16S rRNA gene sequences on the PacBio system. All fermentation parameters, except for NH3-N, showed linear changes with increasing F:C ratios (p ≤ 0.05). F:C ratio affected Isotricha spp. and Dasytricha spp. counts. Ciliate-associated bacterial fractions were significantly less diverse than the total bacterial group, as indicated by richness, phylogenetic diversity, and evenness indices. This suggests potential specific associations within ciliate-provided microhabitats. Both diet and ciliate fractions significantly influenced the overall bacteriome (p ≤ 0.05). More bacteriome features were differentially abundant due to the ciliate fraction effect rather than diet (q ≤ 0.05). Our newly proposed washing procedure, using higher ciliate cell counts and minimal bacterial contamination, effectively removed free-living or loosely associated bacteria. This allows focus on ciliate-associated bacterial populations, which may include potential symbionts or engulfed bacteria of host ruminal ciliates. Verifying these associations could provide insights into rumen microbiome dynamics, nitrogen utilization, hydrogen balance, and microbiome variation under different F:C ratios.https://doi.org/10.1038/s41598-025-10599-6Ruminal ciliateCiliate-associated bacteriaRuminal bacteriomeDietary effectSymbiont
spellingShingle Geonwoo Kim
Woohyung Lee
Tansol Park
Exploring the impact of forage-to-concentrate ratios on the ruminal bacteriome in vitro focusing on ciliate-associated bacteria
Scientific Reports
Ruminal ciliate
Ciliate-associated bacteria
Ruminal bacteriome
Dietary effect
Symbiont
title Exploring the impact of forage-to-concentrate ratios on the ruminal bacteriome in vitro focusing on ciliate-associated bacteria
title_full Exploring the impact of forage-to-concentrate ratios on the ruminal bacteriome in vitro focusing on ciliate-associated bacteria
title_fullStr Exploring the impact of forage-to-concentrate ratios on the ruminal bacteriome in vitro focusing on ciliate-associated bacteria
title_full_unstemmed Exploring the impact of forage-to-concentrate ratios on the ruminal bacteriome in vitro focusing on ciliate-associated bacteria
title_short Exploring the impact of forage-to-concentrate ratios on the ruminal bacteriome in vitro focusing on ciliate-associated bacteria
title_sort exploring the impact of forage to concentrate ratios on the ruminal bacteriome in vitro focusing on ciliate associated bacteria
topic Ruminal ciliate
Ciliate-associated bacteria
Ruminal bacteriome
Dietary effect
Symbiont
url https://doi.org/10.1038/s41598-025-10599-6
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