Bioenergy sorghum stem density increases threefold following internode elongation due to continued accumulation of lignified cell walls and complex regulation of genes involved in cell wall biosynthesis

Abstract Bioenergy sorghum is a highly productive drought tolerant C4 grass that accumulates ~ 80% of its harvested biomass in ~ 4 m long stems comprised of > 40 internodes that develop sequentially during an extended vegetative growth phase. Following elongation of each internode, internode dens...

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Main Authors: Ka Man Jasmine Yu, Brock D. Weers, Brian A. McKinley, Priscilla D. Glenn, Evan Kurtz, William L. Rooney, John E. Mullet
Format: Article
Language:English
Published: BMC 2025-06-01
Series:Biotechnology for Biofuels and Bioproducts
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Online Access:https://doi.org/10.1186/s13068-025-02659-w
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author Ka Man Jasmine Yu
Brock D. Weers
Brian A. McKinley
Priscilla D. Glenn
Evan Kurtz
William L. Rooney
John E. Mullet
author_facet Ka Man Jasmine Yu
Brock D. Weers
Brian A. McKinley
Priscilla D. Glenn
Evan Kurtz
William L. Rooney
John E. Mullet
author_sort Ka Man Jasmine Yu
collection DOAJ
description Abstract Bioenergy sorghum is a highly productive drought tolerant C4 grass that accumulates ~ 80% of its harvested biomass in ~ 4 m long stems comprised of > 40 internodes that develop sequentially during an extended vegetative growth phase. Following elongation of each internode, internode density increases ~ threefold to fourfold primarily due to the accumulation of cell walls composed of cellulose, glucuronoarabinoxylan and lignin. Lignin accumulates initially on cell walls of sclerenchyma cells surrounding vascular bundles and later on cell walls of the stem rind and stem core pith parenchyma. Many genes involved in cell wall biosynthesis were expressed continuously during the stem internode densification process whereas others showed dynamic patterns of expression (high to low, low to high). Several CESA genes involved in primary cell wall cellulose synthesis were expressed in the stem rind and core throughout the stem densification phase. In contrast, CESA genes involved in secondary cell wall biogenesis were expressed continuously in the stem rind but downregulated in the stem core shortly after completion of internode elongation. Overall, accumulation of cell wall biomass in elongated internodes during stem densification increases stem mechanical strength and biomass bulk density while modifying biomass composition in ways that could impact the amount and release of cellulosic sugars and lignin-derived bioproducts.
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spelling doaj-art-efea94c1670b4ed6abfd2bad23da64bb2025-08-20T02:05:45ZengBMCBiotechnology for Biofuels and Bioproducts2731-36542025-06-0118111610.1186/s13068-025-02659-wBioenergy sorghum stem density increases threefold following internode elongation due to continued accumulation of lignified cell walls and complex regulation of genes involved in cell wall biosynthesisKa Man Jasmine Yu0Brock D. Weers1Brian A. McKinley2Priscilla D. Glenn3Evan Kurtz4William L. Rooney5John E. Mullet6Department of Biochemistry and Biophysics, Texas A&M UniversityDepartment of Biochemistry and Biophysics, Texas A&M UniversityDepartment of Biochemistry and Biophysics, Texas A&M UniversityDepartment of Biochemistry and Biophysics, Texas A&M UniversityDepartment of Biochemistry and Biophysics, Texas A&M UniversityDepartment of Soil and Crop Sciences, Texas A&M UniversityDepartment of Biochemistry and Biophysics, Texas A&M UniversityAbstract Bioenergy sorghum is a highly productive drought tolerant C4 grass that accumulates ~ 80% of its harvested biomass in ~ 4 m long stems comprised of > 40 internodes that develop sequentially during an extended vegetative growth phase. Following elongation of each internode, internode density increases ~ threefold to fourfold primarily due to the accumulation of cell walls composed of cellulose, glucuronoarabinoxylan and lignin. Lignin accumulates initially on cell walls of sclerenchyma cells surrounding vascular bundles and later on cell walls of the stem rind and stem core pith parenchyma. Many genes involved in cell wall biosynthesis were expressed continuously during the stem internode densification process whereas others showed dynamic patterns of expression (high to low, low to high). Several CESA genes involved in primary cell wall cellulose synthesis were expressed in the stem rind and core throughout the stem densification phase. In contrast, CESA genes involved in secondary cell wall biogenesis were expressed continuously in the stem rind but downregulated in the stem core shortly after completion of internode elongation. Overall, accumulation of cell wall biomass in elongated internodes during stem densification increases stem mechanical strength and biomass bulk density while modifying biomass composition in ways that could impact the amount and release of cellulosic sugars and lignin-derived bioproducts.https://doi.org/10.1186/s13068-025-02659-wBioenergy sorghumCell wallsStem densityLigninGene expression
spellingShingle Ka Man Jasmine Yu
Brock D. Weers
Brian A. McKinley
Priscilla D. Glenn
Evan Kurtz
William L. Rooney
John E. Mullet
Bioenergy sorghum stem density increases threefold following internode elongation due to continued accumulation of lignified cell walls and complex regulation of genes involved in cell wall biosynthesis
Biotechnology for Biofuels and Bioproducts
Bioenergy sorghum
Cell walls
Stem density
Lignin
Gene expression
title Bioenergy sorghum stem density increases threefold following internode elongation due to continued accumulation of lignified cell walls and complex regulation of genes involved in cell wall biosynthesis
title_full Bioenergy sorghum stem density increases threefold following internode elongation due to continued accumulation of lignified cell walls and complex regulation of genes involved in cell wall biosynthesis
title_fullStr Bioenergy sorghum stem density increases threefold following internode elongation due to continued accumulation of lignified cell walls and complex regulation of genes involved in cell wall biosynthesis
title_full_unstemmed Bioenergy sorghum stem density increases threefold following internode elongation due to continued accumulation of lignified cell walls and complex regulation of genes involved in cell wall biosynthesis
title_short Bioenergy sorghum stem density increases threefold following internode elongation due to continued accumulation of lignified cell walls and complex regulation of genes involved in cell wall biosynthesis
title_sort bioenergy sorghum stem density increases threefold following internode elongation due to continued accumulation of lignified cell walls and complex regulation of genes involved in cell wall biosynthesis
topic Bioenergy sorghum
Cell walls
Stem density
Lignin
Gene expression
url https://doi.org/10.1186/s13068-025-02659-w
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