Species diversity and risk factors of gastrointestinal nematodes in smallholder dairy calves in Kenya

Gastrointestinal nematodes (GIN) are of major concern in dairy farming, particularly in smallholder systems, because of their impact on the health of the calves and later on their productivity. These infections often occur as co-infections, which can complicate their prevention and treatment. The ai...

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Main Authors: Sylvia Cheptoo, Erhan Yalcindag, Lina González Gordon, Benson Rukwaro, Joseph Samuel Kimatu, Joseph Wasonga, Benedict E. Karani, Gideon Ndambuki, Susan Migeni, Jesse Kagai, Linus Eric Kiprotich, Nelson Saya, Deepali Vasoya, Getrude Nangekhe, Justus Onguso, Grace Mungai, Barend Mark Bronsvoort, Elizabeth Anne Jessie Cook
Format: Article
Language:English
Published: Frontiers Media S.A. 2025-08-01
Series:Frontiers in Veterinary Science
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Online Access:https://www.frontiersin.org/articles/10.3389/fvets.2025.1588350/full
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author Sylvia Cheptoo
Sylvia Cheptoo
Erhan Yalcindag
Lina González Gordon
Benson Rukwaro
Benson Rukwaro
Joseph Samuel Kimatu
Joseph Samuel Kimatu
Joseph Wasonga
Joseph Wasonga
Benedict E. Karani
Gideon Ndambuki
Susan Migeni
Jesse Kagai
Linus Eric Kiprotich
Nelson Saya
Deepali Vasoya
Getrude Nangekhe
Justus Onguso
Grace Mungai
Barend Mark Bronsvoort
Elizabeth Anne Jessie Cook
author_facet Sylvia Cheptoo
Sylvia Cheptoo
Erhan Yalcindag
Lina González Gordon
Benson Rukwaro
Benson Rukwaro
Joseph Samuel Kimatu
Joseph Samuel Kimatu
Joseph Wasonga
Joseph Wasonga
Benedict E. Karani
Gideon Ndambuki
Susan Migeni
Jesse Kagai
Linus Eric Kiprotich
Nelson Saya
Deepali Vasoya
Getrude Nangekhe
Justus Onguso
Grace Mungai
Barend Mark Bronsvoort
Elizabeth Anne Jessie Cook
author_sort Sylvia Cheptoo
collection DOAJ
description Gastrointestinal nematodes (GIN) are of major concern in dairy farming, particularly in smallholder systems, because of their impact on the health of the calves and later on their productivity. These infections often occur as co-infections, which can complicate their prevention and treatment. The aim of this study was to conduct fecal egg counts (FEC), genetically identify GIN species, assess species diversity, and identify associated risk factors for GIN infections in dairy calves. Fecal samples were collected from 532 dairy calves across 289 small holder dairy farms. Species identification was achieved through deep amplicon sequencing of the Internal Transcribed Spacer-2 rDNA locus (ITS-2) of first-stage larvae (L1). The mean eggs per gram (EPG) was 62.0 ± 93.0. Most of the calves 64.2% had low-intensity infections (<50 EPG), 28.6% had medium-intensity infections (50–200 EPG), and 7.2% had high-intensity infections (>200 EPG). Next Generation Sequencing analysis identified nine GIN species, with Cooperia punctata (27.8%), Haemonchus placei (26.3%), and Haemonchus contortus (23.6%) being the most prevalent. Co-infections were common, accounting for 69.5% of all infections, with two (40.1%), three (26.9%), and four-species combinations (19.8%) predominating. Male calves showed a significant association with both increased FEC and smaller heart girth, while FEC decreased with age. H. placei and C. punctata were associated with increased FEC, whereas Ostertagia ostertagi (14.5%) and Trichostrongylus colubriformis (8.0%) were associated with decreased heart girth. Calves managed under pasture systems had higher odds of co-infection. This study reveals that GIN infections are highly prevalent in dairy calves, with co-infections being common, and that GIN burden is significantly influenced by calf age, sex, and management system. The Nemabiome tool offers a promising approach to assessing GIN burden and guiding the selection of anthelmintic protocols as part of sustainable farming strategies in tropical regions.
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spelling doaj-art-ee52cb850eee46d7a394c42252273f502025-08-20T03:04:02ZengFrontiers Media S.A.Frontiers in Veterinary Science2297-17692025-08-011210.3389/fvets.2025.15883501588350Species diversity and risk factors of gastrointestinal nematodes in smallholder dairy calves in KenyaSylvia Cheptoo0Sylvia Cheptoo1Erhan Yalcindag2Lina González Gordon3Benson Rukwaro4Benson Rukwaro5Joseph Samuel Kimatu6Joseph Samuel Kimatu7Joseph Wasonga8Joseph Wasonga9Benedict E. Karani10Gideon Ndambuki11Susan Migeni12Jesse Kagai13Linus Eric Kiprotich14Nelson Saya15Deepali Vasoya16Getrude Nangekhe17Justus Onguso18Grace Mungai19Barend Mark Bronsvoort20Elizabeth Anne Jessie Cook21Jomo Kenyatta University of Agriculture and Technology (JKUAT), Nairobi, KenyaCentre for Tropical Livestock Genetics and Health (CTLGH), International Livestock Research Institute (ILRI), Nairobi, KenyaCentre for Tropical Livestock Genetics and Health (CTLGH), Roslin Institute, University of Edinburgh, Easter Bush Campus, Edinburgh, United KingdomCentre for Tropical Livestock Genetics and Health (CTLGH), Roslin Institute, University of Edinburgh, Easter Bush Campus, Edinburgh, United KingdomCentre for Tropical Livestock Genetics and Health (CTLGH), International Livestock Research Institute (ILRI), Nairobi, KenyaDepartment of Veterinary Pathology, Microbiology and Parasitology, The University of Nairobi, Nairobi, KenyaCentre for Tropical Livestock Genetics and Health (CTLGH), International Livestock Research Institute (ILRI), Nairobi, KenyaDepartment of Public Health, Pharmacology and Toxicology, The University of Nairobi, Nairobi, KenyaCentre for Tropical Livestock Genetics and Health (CTLGH), International Livestock Research Institute (ILRI), Nairobi, KenyaDepartment of Community Health, Amref International University (AMIU), Nairobi, KenyaCentre for Tropical Livestock Genetics and Health (CTLGH), International Livestock Research Institute (ILRI), Nairobi, KenyaCentre for Tropical Livestock Genetics and Health (CTLGH), International Livestock Research Institute (ILRI), Nairobi, KenyaCentre for Tropical Livestock Genetics and Health (CTLGH), International Livestock Research Institute (ILRI), Nairobi, KenyaMazingira Centre, International Livestock Research Institute (ILRI), Nairobi, KenyaMazingira Centre, International Livestock Research Institute (ILRI), Nairobi, KenyaMazingira Centre, International Livestock Research Institute (ILRI), Nairobi, KenyaUK Dementia Research Institute at University of Edinburgh, Edinburgh, United KingdomCentre for Tropical Livestock Genetics and Health (CTLGH), International Livestock Research Institute (ILRI), Nairobi, KenyaJomo Kenyatta University of Agriculture and Technology (JKUAT), Nairobi, KenyaJomo Kenyatta University of Agriculture and Technology (JKUAT), Nairobi, KenyaCentre for Tropical Livestock Genetics and Health (CTLGH), Roslin Institute, University of Edinburgh, Easter Bush Campus, Edinburgh, United KingdomCentre for Tropical Livestock Genetics and Health (CTLGH), International Livestock Research Institute (ILRI), Nairobi, KenyaGastrointestinal nematodes (GIN) are of major concern in dairy farming, particularly in smallholder systems, because of their impact on the health of the calves and later on their productivity. These infections often occur as co-infections, which can complicate their prevention and treatment. The aim of this study was to conduct fecal egg counts (FEC), genetically identify GIN species, assess species diversity, and identify associated risk factors for GIN infections in dairy calves. Fecal samples were collected from 532 dairy calves across 289 small holder dairy farms. Species identification was achieved through deep amplicon sequencing of the Internal Transcribed Spacer-2 rDNA locus (ITS-2) of first-stage larvae (L1). The mean eggs per gram (EPG) was 62.0 ± 93.0. Most of the calves 64.2% had low-intensity infections (<50 EPG), 28.6% had medium-intensity infections (50–200 EPG), and 7.2% had high-intensity infections (>200 EPG). Next Generation Sequencing analysis identified nine GIN species, with Cooperia punctata (27.8%), Haemonchus placei (26.3%), and Haemonchus contortus (23.6%) being the most prevalent. Co-infections were common, accounting for 69.5% of all infections, with two (40.1%), three (26.9%), and four-species combinations (19.8%) predominating. Male calves showed a significant association with both increased FEC and smaller heart girth, while FEC decreased with age. H. placei and C. punctata were associated with increased FEC, whereas Ostertagia ostertagi (14.5%) and Trichostrongylus colubriformis (8.0%) were associated with decreased heart girth. Calves managed under pasture systems had higher odds of co-infection. This study reveals that GIN infections are highly prevalent in dairy calves, with co-infections being common, and that GIN burden is significantly influenced by calf age, sex, and management system. The Nemabiome tool offers a promising approach to assessing GIN burden and guiding the selection of anthelmintic protocols as part of sustainable farming strategies in tropical regions.https://www.frontiersin.org/articles/10.3389/fvets.2025.1588350/fullgastrointestinal nematodesdeep amplicon sequencingrisk factorco-infectionsdairy calvesKenya
spellingShingle Sylvia Cheptoo
Sylvia Cheptoo
Erhan Yalcindag
Lina González Gordon
Benson Rukwaro
Benson Rukwaro
Joseph Samuel Kimatu
Joseph Samuel Kimatu
Joseph Wasonga
Joseph Wasonga
Benedict E. Karani
Gideon Ndambuki
Susan Migeni
Jesse Kagai
Linus Eric Kiprotich
Nelson Saya
Deepali Vasoya
Getrude Nangekhe
Justus Onguso
Grace Mungai
Barend Mark Bronsvoort
Elizabeth Anne Jessie Cook
Species diversity and risk factors of gastrointestinal nematodes in smallholder dairy calves in Kenya
Frontiers in Veterinary Science
gastrointestinal nematodes
deep amplicon sequencing
risk factor
co-infections
dairy calves
Kenya
title Species diversity and risk factors of gastrointestinal nematodes in smallholder dairy calves in Kenya
title_full Species diversity and risk factors of gastrointestinal nematodes in smallholder dairy calves in Kenya
title_fullStr Species diversity and risk factors of gastrointestinal nematodes in smallholder dairy calves in Kenya
title_full_unstemmed Species diversity and risk factors of gastrointestinal nematodes in smallholder dairy calves in Kenya
title_short Species diversity and risk factors of gastrointestinal nematodes in smallholder dairy calves in Kenya
title_sort species diversity and risk factors of gastrointestinal nematodes in smallholder dairy calves in kenya
topic gastrointestinal nematodes
deep amplicon sequencing
risk factor
co-infections
dairy calves
Kenya
url https://www.frontiersin.org/articles/10.3389/fvets.2025.1588350/full
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