Critical role of alpha spectrin in DNA repair: the importance of μ-calpain and Fanconi anemia proteins

Nonerythroid spectrins are proteins important in maintaining the structural integrity and flexibility of the cell and nuclear membranes and are essential for a number of functionally important cellular processes. One of these proteins, nonerythroid α spectrin (αSpII), plays a critical role in DNA re...

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Main Author: Muriel W. Lambert
Format: Article
Language:English
Published: Frontiers Media S.A. 2025-05-01
Series:Experimental Biology and Medicine
Subjects:
Online Access:https://www.ebm-journal.org/articles/10.3389/ebm.2025.10537/full
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author Muriel W. Lambert
author_facet Muriel W. Lambert
author_sort Muriel W. Lambert
collection DOAJ
description Nonerythroid spectrins are proteins important in maintaining the structural integrity and flexibility of the cell and nuclear membranes and are essential for a number of functionally important cellular processes. One of these proteins, nonerythroid α spectrin (αSpII), plays a critical role in DNA repair, specifically repair of DNA interstrand crosslinks (ICLs), where it acts as a scaffold, recruiting repair proteins to sites of damage. Loss or breakdown of αSpII is an important factor in a number of disorders. One of these is Fanconi anemia (FA), a genetic disorder characterized by bone marrow failure, chromosome instability, cancer predisposition, congenital abnormalities and a defect in DNA ICL repair. Significantly, breakdown of αSpII occurs in cells from a number of FA complementation groups, due to excessive cleavage by the protease, μ-calpain, leading to defective repair of DNA ICLs in telomeric and non-telomeric DNA. Knockdown of μ-calpain in FA cells by μ-calpain siRNA results in restoration of αSpII levels to normal and repair of DNA ICLs in telomeric and non-telomeric DNA, demonstrating the importance of αSpII stability in the repair process. It is hypothesized that there is a mechanistic link between excessive cleavage of αSpII by μ-calpain and defective DNA ICL repair in FA and that FA proteins, which are deficient in FA, play a key role in maintaining the stability of αSpII and preventing its cleavage by μ-calpain. All of these events are proposed to be important key factors involved in the pathophysiology of FA and suggest new avenues for potential therapeutic intervention.
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spelling doaj-art-ed89a343c2304980b48ea13bb0f3e02d2025-08-20T01:48:26ZengFrontiers Media S.A.Experimental Biology and Medicine1535-36992025-05-0125010.3389/ebm.2025.1053710537Critical role of alpha spectrin in DNA repair: the importance of μ-calpain and Fanconi anemia proteinsMuriel W. LambertNonerythroid spectrins are proteins important in maintaining the structural integrity and flexibility of the cell and nuclear membranes and are essential for a number of functionally important cellular processes. One of these proteins, nonerythroid α spectrin (αSpII), plays a critical role in DNA repair, specifically repair of DNA interstrand crosslinks (ICLs), where it acts as a scaffold, recruiting repair proteins to sites of damage. Loss or breakdown of αSpII is an important factor in a number of disorders. One of these is Fanconi anemia (FA), a genetic disorder characterized by bone marrow failure, chromosome instability, cancer predisposition, congenital abnormalities and a defect in DNA ICL repair. Significantly, breakdown of αSpII occurs in cells from a number of FA complementation groups, due to excessive cleavage by the protease, μ-calpain, leading to defective repair of DNA ICLs in telomeric and non-telomeric DNA. Knockdown of μ-calpain in FA cells by μ-calpain siRNA results in restoration of αSpII levels to normal and repair of DNA ICLs in telomeric and non-telomeric DNA, demonstrating the importance of αSpII stability in the repair process. It is hypothesized that there is a mechanistic link between excessive cleavage of αSpII by μ-calpain and defective DNA ICL repair in FA and that FA proteins, which are deficient in FA, play a key role in maintaining the stability of αSpII and preventing its cleavage by μ-calpain. All of these events are proposed to be important key factors involved in the pathophysiology of FA and suggest new avenues for potential therapeutic intervention.https://www.ebm-journal.org/articles/10.3389/ebm.2025.10537/fullalpha spectrinDNA repairDNA interstrand crosslinksμ-calpainFanconi anemia proteins
spellingShingle Muriel W. Lambert
Critical role of alpha spectrin in DNA repair: the importance of μ-calpain and Fanconi anemia proteins
Experimental Biology and Medicine
alpha spectrin
DNA repair
DNA interstrand crosslinks
μ-calpain
Fanconi anemia proteins
title Critical role of alpha spectrin in DNA repair: the importance of μ-calpain and Fanconi anemia proteins
title_full Critical role of alpha spectrin in DNA repair: the importance of μ-calpain and Fanconi anemia proteins
title_fullStr Critical role of alpha spectrin in DNA repair: the importance of μ-calpain and Fanconi anemia proteins
title_full_unstemmed Critical role of alpha spectrin in DNA repair: the importance of μ-calpain and Fanconi anemia proteins
title_short Critical role of alpha spectrin in DNA repair: the importance of μ-calpain and Fanconi anemia proteins
title_sort critical role of alpha spectrin in dna repair the importance of μ calpain and fanconi anemia proteins
topic alpha spectrin
DNA repair
DNA interstrand crosslinks
μ-calpain
Fanconi anemia proteins
url https://www.ebm-journal.org/articles/10.3389/ebm.2025.10537/full
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