The architecture of gene regulatory variation across multiple human tissues: the MuTHER study.

While there have been studies exploring regulatory variation in one or more tissues, the complexity of tissue-specificity in multiple primary tissues is not yet well understood. We explore in depth the role of cis-regulatory variation in three human tissues: lymphoblastoid cell lines (LCL), skin, an...

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Main Authors: Alexandra C Nica, Leopold Parts, Daniel Glass, James Nisbet, Amy Barrett, Magdalena Sekowska, Mary Travers, Simon Potter, Elin Grundberg, Kerrin Small, Asa K Hedman, Veronique Bataille, Jordana Tzenova Bell, Gabriela Surdulescu, Antigone S Dimas, Catherine Ingle, Frank O Nestle, Paola di Meglio, Josine L Min, Alicja Wilk, Christopher J Hammond, Neelam Hassanali, Tsun-Po Yang, Stephen B Montgomery, Steve O'Rahilly, Cecilia M Lindgren, Krina T Zondervan, Nicole Soranzo, Inês Barroso, Richard Durbin, Kourosh Ahmadi, Panos Deloukas, Mark I McCarthy, Emmanouil T Dermitzakis, Timothy D Spector, MuTHER Consortium
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2011-02-01
Series:PLoS Genetics
Online Access:https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1002003&type=printable
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author Alexandra C Nica
Leopold Parts
Daniel Glass
James Nisbet
Amy Barrett
Magdalena Sekowska
Mary Travers
Simon Potter
Elin Grundberg
Kerrin Small
Asa K Hedman
Veronique Bataille
Jordana Tzenova Bell
Gabriela Surdulescu
Antigone S Dimas
Catherine Ingle
Frank O Nestle
Paola di Meglio
Josine L Min
Alicja Wilk
Christopher J Hammond
Neelam Hassanali
Tsun-Po Yang
Stephen B Montgomery
Steve O'Rahilly
Cecilia M Lindgren
Krina T Zondervan
Nicole Soranzo
Inês Barroso
Richard Durbin
Kourosh Ahmadi
Panos Deloukas
Mark I McCarthy
Emmanouil T Dermitzakis
Timothy D Spector
MuTHER Consortium
author_facet Alexandra C Nica
Leopold Parts
Daniel Glass
James Nisbet
Amy Barrett
Magdalena Sekowska
Mary Travers
Simon Potter
Elin Grundberg
Kerrin Small
Asa K Hedman
Veronique Bataille
Jordana Tzenova Bell
Gabriela Surdulescu
Antigone S Dimas
Catherine Ingle
Frank O Nestle
Paola di Meglio
Josine L Min
Alicja Wilk
Christopher J Hammond
Neelam Hassanali
Tsun-Po Yang
Stephen B Montgomery
Steve O'Rahilly
Cecilia M Lindgren
Krina T Zondervan
Nicole Soranzo
Inês Barroso
Richard Durbin
Kourosh Ahmadi
Panos Deloukas
Mark I McCarthy
Emmanouil T Dermitzakis
Timothy D Spector
MuTHER Consortium
author_sort Alexandra C Nica
collection DOAJ
description While there have been studies exploring regulatory variation in one or more tissues, the complexity of tissue-specificity in multiple primary tissues is not yet well understood. We explore in depth the role of cis-regulatory variation in three human tissues: lymphoblastoid cell lines (LCL), skin, and fat. The samples (156 LCL, 160 skin, 166 fat) were derived simultaneously from a subset of well-phenotyped healthy female twins of the MuTHER resource. We discover an abundance of cis-eQTLs in each tissue similar to previous estimates (858 or 4.7% of genes). In addition, we apply factor analysis (FA) to remove effects of latent variables, thus more than doubling the number of our discoveries (1,822 eQTL genes). The unique study design (Matched Co-Twin Analysis--MCTA) permits immediate replication of eQTLs using co-twins (93%-98%) and validation of the considerable gain in eQTL discovery after FA correction. We highlight the challenges of comparing eQTLs between tissues. After verifying previous significance threshold-based estimates of tissue-specificity, we show their limitations given their dependency on statistical power. We propose that continuous estimates of the proportion of tissue-shared signals and direct comparison of the magnitude of effect on the fold change in expression are essential properties that jointly provide a biologically realistic view of tissue-specificity. Under this framework we demonstrate that 30% of eQTLs are shared among the three tissues studied, while another 29% appear exclusively tissue-specific. However, even among the shared eQTLs, a substantial proportion (10%-20%) have significant differences in the magnitude of fold change between genotypic classes across tissues. Our results underline the need to account for the complexity of eQTL tissue-specificity in an effort to assess consequences of such variants for complex traits.
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spelling doaj-art-ed6d41d2e4874ff2b5d076c62e716ffe2025-08-20T02:08:57ZengPublic Library of Science (PLoS)PLoS Genetics1553-73901553-74042011-02-0172e100200310.1371/journal.pgen.1002003The architecture of gene regulatory variation across multiple human tissues: the MuTHER study.Alexandra C NicaLeopold PartsDaniel GlassJames NisbetAmy BarrettMagdalena SekowskaMary TraversSimon PotterElin GrundbergKerrin SmallAsa K HedmanVeronique BatailleJordana Tzenova BellGabriela SurdulescuAntigone S DimasCatherine IngleFrank O NestlePaola di MeglioJosine L MinAlicja WilkChristopher J HammondNeelam HassanaliTsun-Po YangStephen B MontgomerySteve O'RahillyCecilia M LindgrenKrina T ZondervanNicole SoranzoInês BarrosoRichard DurbinKourosh AhmadiPanos DeloukasMark I McCarthyEmmanouil T DermitzakisTimothy D SpectorMuTHER ConsortiumWhile there have been studies exploring regulatory variation in one or more tissues, the complexity of tissue-specificity in multiple primary tissues is not yet well understood. We explore in depth the role of cis-regulatory variation in three human tissues: lymphoblastoid cell lines (LCL), skin, and fat. The samples (156 LCL, 160 skin, 166 fat) were derived simultaneously from a subset of well-phenotyped healthy female twins of the MuTHER resource. We discover an abundance of cis-eQTLs in each tissue similar to previous estimates (858 or 4.7% of genes). In addition, we apply factor analysis (FA) to remove effects of latent variables, thus more than doubling the number of our discoveries (1,822 eQTL genes). The unique study design (Matched Co-Twin Analysis--MCTA) permits immediate replication of eQTLs using co-twins (93%-98%) and validation of the considerable gain in eQTL discovery after FA correction. We highlight the challenges of comparing eQTLs between tissues. After verifying previous significance threshold-based estimates of tissue-specificity, we show their limitations given their dependency on statistical power. We propose that continuous estimates of the proportion of tissue-shared signals and direct comparison of the magnitude of effect on the fold change in expression are essential properties that jointly provide a biologically realistic view of tissue-specificity. Under this framework we demonstrate that 30% of eQTLs are shared among the three tissues studied, while another 29% appear exclusively tissue-specific. However, even among the shared eQTLs, a substantial proportion (10%-20%) have significant differences in the magnitude of fold change between genotypic classes across tissues. Our results underline the need to account for the complexity of eQTL tissue-specificity in an effort to assess consequences of such variants for complex traits.https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1002003&type=printable
spellingShingle Alexandra C Nica
Leopold Parts
Daniel Glass
James Nisbet
Amy Barrett
Magdalena Sekowska
Mary Travers
Simon Potter
Elin Grundberg
Kerrin Small
Asa K Hedman
Veronique Bataille
Jordana Tzenova Bell
Gabriela Surdulescu
Antigone S Dimas
Catherine Ingle
Frank O Nestle
Paola di Meglio
Josine L Min
Alicja Wilk
Christopher J Hammond
Neelam Hassanali
Tsun-Po Yang
Stephen B Montgomery
Steve O'Rahilly
Cecilia M Lindgren
Krina T Zondervan
Nicole Soranzo
Inês Barroso
Richard Durbin
Kourosh Ahmadi
Panos Deloukas
Mark I McCarthy
Emmanouil T Dermitzakis
Timothy D Spector
MuTHER Consortium
The architecture of gene regulatory variation across multiple human tissues: the MuTHER study.
PLoS Genetics
title The architecture of gene regulatory variation across multiple human tissues: the MuTHER study.
title_full The architecture of gene regulatory variation across multiple human tissues: the MuTHER study.
title_fullStr The architecture of gene regulatory variation across multiple human tissues: the MuTHER study.
title_full_unstemmed The architecture of gene regulatory variation across multiple human tissues: the MuTHER study.
title_short The architecture of gene regulatory variation across multiple human tissues: the MuTHER study.
title_sort architecture of gene regulatory variation across multiple human tissues the muther study
url https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1002003&type=printable
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