OsPIP2;4 aquaporin water channel primarily expressed in roots of rice mediates both water and nonselective Na+ and K+ conductance

Abstract Aquaporin (AQP)-dependent water transport across membranes is indispensable in plants. Recent evidence shows that several AQPs, including plasma membrane intrinsic proteins (PIPs), facilitate the electrogenic transport of ions as well as water transport and are referred to as ion-conducting...

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Main Authors: Sen Thi Huong Tran, Maki Katsuhara, Yunosuke Mito, Aya Onishi, Ayaka Higa, Shuntaro Ono, Newton Chandra Paul, Rie Horie, Yoshihiko Harada, Tomoaki Horie
Format: Article
Language:English
Published: Nature Portfolio 2025-04-01
Series:Scientific Reports
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Online Access:https://doi.org/10.1038/s41598-025-96259-1
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author Sen Thi Huong Tran
Maki Katsuhara
Yunosuke Mito
Aya Onishi
Ayaka Higa
Shuntaro Ono
Newton Chandra Paul
Rie Horie
Yoshihiko Harada
Tomoaki Horie
author_facet Sen Thi Huong Tran
Maki Katsuhara
Yunosuke Mito
Aya Onishi
Ayaka Higa
Shuntaro Ono
Newton Chandra Paul
Rie Horie
Yoshihiko Harada
Tomoaki Horie
author_sort Sen Thi Huong Tran
collection DOAJ
description Abstract Aquaporin (AQP)-dependent water transport across membranes is indispensable in plants. Recent evidence shows that several AQPs, including plasma membrane intrinsic proteins (PIPs), facilitate the electrogenic transport of ions as well as water transport and are referred to as ion-conducting aquaporins (icAQPs). The present study attempted to identify icAQPs that exhibit cation transport activity among PIPs from rice. Electrophysiological experiments on 11 OsPIPs using Xenopus laevis oocytes revealed that OsPIP2;4 mediated the electrogenic transport of alkali monovalent cations with the selectivity sequence of Na+ ≈ K+ > Rb+ > Cs+ > Li+, suggesting non-selective cation conductance for Na+ and K+. Transcripts of OsPIP2;4 were abundant in the elongation and mature zones of roots with similar expression levels between the root stelar and remaining outer parts in the cultivar Nipponbare. Immunostaining using sections of the crown roots of Nipponbare plants revealed the expression of OsPIP2;4 in the exodermis and sclerenchyma of the surface region and in the endodermis and pericycle of the stelar region. The present results provide novel insights into OsPIP2;4-dependent non-selective Na+ and K+ transport and its physiological roles in rice.
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spelling doaj-art-ec3d3a53a2e448f8b7c2924e57e627502025-08-20T02:24:29ZengNature PortfolioScientific Reports2045-23222025-04-0115111210.1038/s41598-025-96259-1OsPIP2;4 aquaporin water channel primarily expressed in roots of rice mediates both water and nonselective Na+ and K+ conductanceSen Thi Huong Tran0Maki Katsuhara1Yunosuke Mito2Aya Onishi3Ayaka Higa4Shuntaro Ono5Newton Chandra Paul6Rie Horie7Yoshihiko Harada8Tomoaki Horie9Institute of Plant Science and Resources, Okayama UniversityInstitute of Plant Science and Resources, Okayama UniversityDivision of Applied Biology, Faculty of Textile Science and Technology, Shinshu UniversityInstitute of Plant Science and Resources, Okayama UniversityDivision of Applied Biology, Faculty of Textile Science and Technology, Shinshu UniversityInstitute of Plant Science and Resources, Okayama UniversityInstitute of Plant Science and Resources, Okayama UniversityDivision of Applied Biology, Faculty of Textile Science and Technology, Shinshu UniversityDivision of Applied Biology, Faculty of Textile Science and Technology, Shinshu UniversityDivision of Applied Biology, Faculty of Textile Science and Technology, Shinshu UniversityAbstract Aquaporin (AQP)-dependent water transport across membranes is indispensable in plants. Recent evidence shows that several AQPs, including plasma membrane intrinsic proteins (PIPs), facilitate the electrogenic transport of ions as well as water transport and are referred to as ion-conducting aquaporins (icAQPs). The present study attempted to identify icAQPs that exhibit cation transport activity among PIPs from rice. Electrophysiological experiments on 11 OsPIPs using Xenopus laevis oocytes revealed that OsPIP2;4 mediated the electrogenic transport of alkali monovalent cations with the selectivity sequence of Na+ ≈ K+ > Rb+ > Cs+ > Li+, suggesting non-selective cation conductance for Na+ and K+. Transcripts of OsPIP2;4 were abundant in the elongation and mature zones of roots with similar expression levels between the root stelar and remaining outer parts in the cultivar Nipponbare. Immunostaining using sections of the crown roots of Nipponbare plants revealed the expression of OsPIP2;4 in the exodermis and sclerenchyma of the surface region and in the endodermis and pericycle of the stelar region. The present results provide novel insights into OsPIP2;4-dependent non-selective Na+ and K+ transport and its physiological roles in rice.https://doi.org/10.1038/s41598-025-96259-1Ion-conducting AquaporinsNon-selective cation channelRiceRoots
spellingShingle Sen Thi Huong Tran
Maki Katsuhara
Yunosuke Mito
Aya Onishi
Ayaka Higa
Shuntaro Ono
Newton Chandra Paul
Rie Horie
Yoshihiko Harada
Tomoaki Horie
OsPIP2;4 aquaporin water channel primarily expressed in roots of rice mediates both water and nonselective Na+ and K+ conductance
Scientific Reports
Ion-conducting Aquaporins
Non-selective cation channel
Rice
Roots
title OsPIP2;4 aquaporin water channel primarily expressed in roots of rice mediates both water and nonselective Na+ and K+ conductance
title_full OsPIP2;4 aquaporin water channel primarily expressed in roots of rice mediates both water and nonselective Na+ and K+ conductance
title_fullStr OsPIP2;4 aquaporin water channel primarily expressed in roots of rice mediates both water and nonselective Na+ and K+ conductance
title_full_unstemmed OsPIP2;4 aquaporin water channel primarily expressed in roots of rice mediates both water and nonselective Na+ and K+ conductance
title_short OsPIP2;4 aquaporin water channel primarily expressed in roots of rice mediates both water and nonselective Na+ and K+ conductance
title_sort ospip2 4 aquaporin water channel primarily expressed in roots of rice mediates both water and nonselective na and k conductance
topic Ion-conducting Aquaporins
Non-selective cation channel
Rice
Roots
url https://doi.org/10.1038/s41598-025-96259-1
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