Comparative analysis of Diospyros (Ebenaceae) plastomes: Insights into genomic features, mutational hotspots, and adaptive evolution

Abstract Diospyros (Ebenaceae) is a widely distributed genus of trees and shrubs from pantropical to temperate regions, with numerous species valued for their fruits (persimmons), timber, and medicinal values. However, information regarding their plastomes and chloroplast evolution is scarce. The pr...

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Main Authors: Yue Huang, Qing Ma, Jing Sun, Li‐Na Zhou, Chan‐Juan Lai, Pan Li, Xin‐Jie Jin, Yong‐Hua Zhang
Format: Article
Language:English
Published: Wiley 2023-07-01
Series:Ecology and Evolution
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Online Access:https://doi.org/10.1002/ece3.10301
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author Yue Huang
Qing Ma
Jing Sun
Li‐Na Zhou
Chan‐Juan Lai
Pan Li
Xin‐Jie Jin
Yong‐Hua Zhang
author_facet Yue Huang
Qing Ma
Jing Sun
Li‐Na Zhou
Chan‐Juan Lai
Pan Li
Xin‐Jie Jin
Yong‐Hua Zhang
author_sort Yue Huang
collection DOAJ
description Abstract Diospyros (Ebenaceae) is a widely distributed genus of trees and shrubs from pantropical to temperate regions, with numerous species valued for their fruits (persimmons), timber, and medicinal values. However, information regarding their plastomes and chloroplast evolution is scarce. The present study performed comparative genomic and evolutionary analyses on plastomes of 45 accepted Diospyros species, including three newly sequenced ones. Our study showed a highly conserved genomic structure across the Diospyros species, with 135–136 encoding genes, including 89 protein‐coding genes, 1–2 pseudogenes (Ψycf1 for all, Ψrps19 for a few), 37 tRNA genes and 8 rRNA genes. Comparative analysis of Diospyros identified three intergenic regions (ccsA‐ndhD, rps16‐psbK and petA‐psbJ) and five genes (rpl33, rpl22, petL, psaC and rps15) as the mutational hotspots in these species. Phylogenomic analysis identified the phylogenetic position of three newly sequenced ones and well supported a monophylogenetic (sub)temperate taxa and four clades in the pantropical taxa. The analysis codon usage identified 30 codons with relative synonymous codon usage (RSCU) values >1 and 29 codons ending with A and U bases. A total of three codons (UUA, GCU, and AGA) with highest RSCU values were identified as the optimal codons. Effective number of codons (ENC)‐plot indicated the significant role of mutational pressure in shaping codon usage, while most protein‐coding genes in Diospyros experienced relaxed purifying selection (dN/dS < 1). Additionally, the psbH gene showed positive selection (dN/dS > 1) in the (sub)temperate species. Thus, the results provide a meaningful foundation for further elaborating Diospyros's genetic architecture and taxonomy, enriching genetic diversity and conserving genetic resources.
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spelling doaj-art-ebe430f95633466b9786f5d0d07a644a2025-08-20T03:01:35ZengWileyEcology and Evolution2045-77582023-07-01137n/an/a10.1002/ece3.10301Comparative analysis of Diospyros (Ebenaceae) plastomes: Insights into genomic features, mutational hotspots, and adaptive evolutionYue Huang0Qing Ma1Jing Sun2Li‐Na Zhou3Chan‐Juan Lai4Pan Li5Xin‐Jie Jin6Yong‐Hua Zhang7College of Life and Environmental Science Wenzhou University Wenzhou ChinaCollege of Biology and Environmental Engineering Zhejiang Shuren University Hangzhou ChinaCollege of Life and Environmental Science Wenzhou University Wenzhou ChinaCollege of Life and Environmental Science Wenzhou University Wenzhou ChinaCollege of Life and Environmental Science Wenzhou University Wenzhou ChinaLaboratory of Systematic & Evolutionary Botany and Biodiversity, College of Life Sciences Zhejiang University Hangzhou ChinaCollege of Life and Environmental Science Wenzhou University Wenzhou ChinaCollege of Life and Environmental Science Wenzhou University Wenzhou ChinaAbstract Diospyros (Ebenaceae) is a widely distributed genus of trees and shrubs from pantropical to temperate regions, with numerous species valued for their fruits (persimmons), timber, and medicinal values. However, information regarding their plastomes and chloroplast evolution is scarce. The present study performed comparative genomic and evolutionary analyses on plastomes of 45 accepted Diospyros species, including three newly sequenced ones. Our study showed a highly conserved genomic structure across the Diospyros species, with 135–136 encoding genes, including 89 protein‐coding genes, 1–2 pseudogenes (Ψycf1 for all, Ψrps19 for a few), 37 tRNA genes and 8 rRNA genes. Comparative analysis of Diospyros identified three intergenic regions (ccsA‐ndhD, rps16‐psbK and petA‐psbJ) and five genes (rpl33, rpl22, petL, psaC and rps15) as the mutational hotspots in these species. Phylogenomic analysis identified the phylogenetic position of three newly sequenced ones and well supported a monophylogenetic (sub)temperate taxa and four clades in the pantropical taxa. The analysis codon usage identified 30 codons with relative synonymous codon usage (RSCU) values >1 and 29 codons ending with A and U bases. A total of three codons (UUA, GCU, and AGA) with highest RSCU values were identified as the optimal codons. Effective number of codons (ENC)‐plot indicated the significant role of mutational pressure in shaping codon usage, while most protein‐coding genes in Diospyros experienced relaxed purifying selection (dN/dS < 1). Additionally, the psbH gene showed positive selection (dN/dS > 1) in the (sub)temperate species. Thus, the results provide a meaningful foundation for further elaborating Diospyros's genetic architecture and taxonomy, enriching genetic diversity and conserving genetic resources.https://doi.org/10.1002/ece3.10301Diospyrosgenetic diversityhyper‐variable regionplastome
spellingShingle Yue Huang
Qing Ma
Jing Sun
Li‐Na Zhou
Chan‐Juan Lai
Pan Li
Xin‐Jie Jin
Yong‐Hua Zhang
Comparative analysis of Diospyros (Ebenaceae) plastomes: Insights into genomic features, mutational hotspots, and adaptive evolution
Ecology and Evolution
Diospyros
genetic diversity
hyper‐variable region
plastome
title Comparative analysis of Diospyros (Ebenaceae) plastomes: Insights into genomic features, mutational hotspots, and adaptive evolution
title_full Comparative analysis of Diospyros (Ebenaceae) plastomes: Insights into genomic features, mutational hotspots, and adaptive evolution
title_fullStr Comparative analysis of Diospyros (Ebenaceae) plastomes: Insights into genomic features, mutational hotspots, and adaptive evolution
title_full_unstemmed Comparative analysis of Diospyros (Ebenaceae) plastomes: Insights into genomic features, mutational hotspots, and adaptive evolution
title_short Comparative analysis of Diospyros (Ebenaceae) plastomes: Insights into genomic features, mutational hotspots, and adaptive evolution
title_sort comparative analysis of diospyros ebenaceae plastomes insights into genomic features mutational hotspots and adaptive evolution
topic Diospyros
genetic diversity
hyper‐variable region
plastome
url https://doi.org/10.1002/ece3.10301
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