Plantar flexors are the main engine of walking in healthy adults

IntroductionThe plantar flexors contribute to the uniqueness of man's walking across bipeds (including apes). This role is achieved in late infancy through neural maturation. This may explain why this mechanism is lost with all corticospinal lesions despite the spared power of plantar flexors i...

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Main Authors: Viviana Rota, Antonio Caronni, Stefano Scarano, Maurizio Amadei, Luigi Tesio
Format: Article
Language:English
Published: Frontiers Media S.A. 2025-07-01
Series:Frontiers in Sports and Active Living
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Online Access:https://www.frontiersin.org/articles/10.3389/fspor.2025.1595065/full
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author Viviana Rota
Antonio Caronni
Antonio Caronni
Stefano Scarano
Maurizio Amadei
Luigi Tesio
author_facet Viviana Rota
Antonio Caronni
Antonio Caronni
Stefano Scarano
Maurizio Amadei
Luigi Tesio
author_sort Viviana Rota
collection DOAJ
description IntroductionThe plantar flexors contribute to the uniqueness of man's walking across bipeds (including apes). This role is achieved in late infancy through neural maturation. This may explain why this mechanism is lost with all corticospinal lesions despite the spared power of plantar flexors in segmental motions. During adult human walking, the plantar flexor muscles at the rear limb, during double stance, are suspected to provide most of the work and power required to translate the body system, which can be represented mechanically by its centre of mass (CoM). However, direct evidence of the dominant role of the ankle muscles in CoM translation is scarce. Experimental evidence requires synchronously assessing the lower limb joints’ and CoM's power.MethodsIn this work, ten healthy adults were requested to walk on a split-belt force treadmill at speeds ranging from 0.3 to 1.2 m s−1. A series of eight subsequent strides was analysed at each different speed. The synchronous analysis of ground reaction forces (through force platforms) and joint rotations (through an optoelectronic system) allowed us to simultaneously measure the CoM and the lower limb joints’ power.ResultsThe dominant role of the ankle plantar flexors, suggested by previous studies focusing on speeds above 0.9 m s−1, was confirmed by observing that changes in ankle power during the push-off phase (end of single stance and initial double stance) mirror the changes in power of the CoM. In the double support phase, the amplitude of the increments in ankle joint power was a strong predictor of the increments in CoM power (R2 = 82%).DiscussionLow walking speeds have been included to foster the interpretation of pathologic gaits, and clinical correlates of these findings in motor impairments are highlighted.Clinical Trial RegistrationClinicalTrials.gov, identifier NCT05778474.
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spelling doaj-art-e97d8d97941d4132b3409420c1fc78ea2025-08-20T03:33:22ZengFrontiers Media S.A.Frontiers in Sports and Active Living2624-93672025-07-01710.3389/fspor.2025.15950651595065Plantar flexors are the main engine of walking in healthy adultsViviana Rota0Antonio Caronni1Antonio Caronni2Stefano Scarano3Maurizio Amadei4Luigi Tesio5IRCCS Istituto Auxologico Italiano, Department of Neurorehabilitation Sciences, Ospedale San Luca, Milan, ItalyIRCCS Istituto Auxologico Italiano, Department of Neurorehabilitation Sciences, Ospedale San Luca, Milan, ItalyDepartment of Biomedical Sciences for Health, Università Degli Studi di Milano, Milan, ItalyIRCCS Istituto Auxologico Italiano, Department of Neurorehabilitation Sciences, Ospedale San Luca, Milan, ItalyIRCCS Istituto Auxologico Italiano, Department of Neurorehabilitation Sciences, Ospedale San Luca, Milan, ItalyIRCCS Istituto Auxologico Italiano, Department of Neurorehabilitation Sciences, Ospedale San Luca, Milan, ItalyIntroductionThe plantar flexors contribute to the uniqueness of man's walking across bipeds (including apes). This role is achieved in late infancy through neural maturation. This may explain why this mechanism is lost with all corticospinal lesions despite the spared power of plantar flexors in segmental motions. During adult human walking, the plantar flexor muscles at the rear limb, during double stance, are suspected to provide most of the work and power required to translate the body system, which can be represented mechanically by its centre of mass (CoM). However, direct evidence of the dominant role of the ankle muscles in CoM translation is scarce. Experimental evidence requires synchronously assessing the lower limb joints’ and CoM's power.MethodsIn this work, ten healthy adults were requested to walk on a split-belt force treadmill at speeds ranging from 0.3 to 1.2 m s−1. A series of eight subsequent strides was analysed at each different speed. The synchronous analysis of ground reaction forces (through force platforms) and joint rotations (through an optoelectronic system) allowed us to simultaneously measure the CoM and the lower limb joints’ power.ResultsThe dominant role of the ankle plantar flexors, suggested by previous studies focusing on speeds above 0.9 m s−1, was confirmed by observing that changes in ankle power during the push-off phase (end of single stance and initial double stance) mirror the changes in power of the CoM. In the double support phase, the amplitude of the increments in ankle joint power was a strong predictor of the increments in CoM power (R2 = 82%).DiscussionLow walking speeds have been included to foster the interpretation of pathologic gaits, and clinical correlates of these findings in motor impairments are highlighted.Clinical Trial RegistrationClinicalTrials.gov, identifier NCT05778474.https://www.frontiersin.org/articles/10.3389/fspor.2025.1595065/fullwalkingmanmuscle powerfootcentre of massneural maturation
spellingShingle Viviana Rota
Antonio Caronni
Antonio Caronni
Stefano Scarano
Maurizio Amadei
Luigi Tesio
Plantar flexors are the main engine of walking in healthy adults
Frontiers in Sports and Active Living
walking
man
muscle power
foot
centre of mass
neural maturation
title Plantar flexors are the main engine of walking in healthy adults
title_full Plantar flexors are the main engine of walking in healthy adults
title_fullStr Plantar flexors are the main engine of walking in healthy adults
title_full_unstemmed Plantar flexors are the main engine of walking in healthy adults
title_short Plantar flexors are the main engine of walking in healthy adults
title_sort plantar flexors are the main engine of walking in healthy adults
topic walking
man
muscle power
foot
centre of mass
neural maturation
url https://www.frontiersin.org/articles/10.3389/fspor.2025.1595065/full
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