A Yeast Mutant Deleted of GPH1 Bears Defects in Lipid Metabolism.

In a previous study we demonstrated up-regulation of the yeast GPH1 gene under conditions of phosphatidylethanolamine (PE) depletion caused by deletion of the mitochondrial (M) phosphatidylserine decarboxylase 1 (PSD1) (Gsell et al., 2013, PLoS One. 8(10):e77380. doi: 10.1371/journal.pone.0077380)....

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Main Authors: Martina Gsell, Ariane Fankl, Lisa Klug, Gerald Mascher, Claudia Schmidt, Claudia Hrastnik, Günther Zellnig, Günther Daum
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2015-01-01
Series:PLoS ONE
Online Access:https://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0136957&type=printable
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author Martina Gsell
Ariane Fankl
Lisa Klug
Gerald Mascher
Claudia Schmidt
Claudia Hrastnik
Günther Zellnig
Günther Daum
author_facet Martina Gsell
Ariane Fankl
Lisa Klug
Gerald Mascher
Claudia Schmidt
Claudia Hrastnik
Günther Zellnig
Günther Daum
author_sort Martina Gsell
collection DOAJ
description In a previous study we demonstrated up-regulation of the yeast GPH1 gene under conditions of phosphatidylethanolamine (PE) depletion caused by deletion of the mitochondrial (M) phosphatidylserine decarboxylase 1 (PSD1) (Gsell et al., 2013, PLoS One. 8(10):e77380. doi: 10.1371/journal.pone.0077380). Gph1p has originally been identified as a glycogen phosphorylase catalyzing degradation of glycogen to glucose in the stationary growth phase of the yeast. Here we show that deletion of this gene also causes decreased levels of phosphatidylcholine (PC), triacylglycerols and steryl esters. Depletion of the two non-polar lipids in a Δgph1 strain leads to lack of lipid droplets, and decrease of the PC level results in instability of the plasma membrane. In vivo labeling experiments revealed that formation of PC via both pathways of biosynthesis, the cytidine diphosphate (CDP)-choline and the methylation route, is negatively affected by a Δgph1 mutation, although expression of genes involved is not down regulated. Altogether, Gph1p besides its function as a glycogen mobilizing enzyme appears to play a regulatory role in yeast lipid metabolism.
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spelling doaj-art-e8b3f8bb9d3b4e9d9ac7d5a085c4bd332025-08-20T02:22:46ZengPublic Library of Science (PLoS)PLoS ONE1932-62032015-01-01109e013695710.1371/journal.pone.0136957A Yeast Mutant Deleted of GPH1 Bears Defects in Lipid Metabolism.Martina GsellAriane FanklLisa KlugGerald MascherClaudia SchmidtClaudia HrastnikGünther ZellnigGünther DaumIn a previous study we demonstrated up-regulation of the yeast GPH1 gene under conditions of phosphatidylethanolamine (PE) depletion caused by deletion of the mitochondrial (M) phosphatidylserine decarboxylase 1 (PSD1) (Gsell et al., 2013, PLoS One. 8(10):e77380. doi: 10.1371/journal.pone.0077380). Gph1p has originally been identified as a glycogen phosphorylase catalyzing degradation of glycogen to glucose in the stationary growth phase of the yeast. Here we show that deletion of this gene also causes decreased levels of phosphatidylcholine (PC), triacylglycerols and steryl esters. Depletion of the two non-polar lipids in a Δgph1 strain leads to lack of lipid droplets, and decrease of the PC level results in instability of the plasma membrane. In vivo labeling experiments revealed that formation of PC via both pathways of biosynthesis, the cytidine diphosphate (CDP)-choline and the methylation route, is negatively affected by a Δgph1 mutation, although expression of genes involved is not down regulated. Altogether, Gph1p besides its function as a glycogen mobilizing enzyme appears to play a regulatory role in yeast lipid metabolism.https://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0136957&type=printable
spellingShingle Martina Gsell
Ariane Fankl
Lisa Klug
Gerald Mascher
Claudia Schmidt
Claudia Hrastnik
Günther Zellnig
Günther Daum
A Yeast Mutant Deleted of GPH1 Bears Defects in Lipid Metabolism.
PLoS ONE
title A Yeast Mutant Deleted of GPH1 Bears Defects in Lipid Metabolism.
title_full A Yeast Mutant Deleted of GPH1 Bears Defects in Lipid Metabolism.
title_fullStr A Yeast Mutant Deleted of GPH1 Bears Defects in Lipid Metabolism.
title_full_unstemmed A Yeast Mutant Deleted of GPH1 Bears Defects in Lipid Metabolism.
title_short A Yeast Mutant Deleted of GPH1 Bears Defects in Lipid Metabolism.
title_sort yeast mutant deleted of gph1 bears defects in lipid metabolism
url https://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0136957&type=printable
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