The INAVA mRNA in Extracellular Vesicles Activates Normal Ovarian Fibroblasts by Phosphorylation–Ubiquitylation Crosstalk of HMGA2

Abstract Ovarian cancer is an aggressive gynecological tumor usually diagnosed with widespread metastases. Extracellular vesicles (EVs), though recognized as important mediators of tumor metastasis, have received limited attention into their specific functions via the mRNA profiling. Here it is repo...

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Main Authors: Lingkai Gu, Zhangjin Shen, Shizhen Shen, Conghui Wang, Yuwan Liu, Xinyi Wei, Mengxia Zheng, Jiaxin Gu, Xiaojing Chen, Yi Sun, Junfen Xu, Yan Lu, Weiguo Lu
Format: Article
Language:English
Published: Wiley 2025-07-01
Series:Advanced Science
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Online Access:https://doi.org/10.1002/advs.202500912
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author Lingkai Gu
Zhangjin Shen
Shizhen Shen
Conghui Wang
Yuwan Liu
Xinyi Wei
Mengxia Zheng
Jiaxin Gu
Xiaojing Chen
Yi Sun
Junfen Xu
Yan Lu
Weiguo Lu
author_facet Lingkai Gu
Zhangjin Shen
Shizhen Shen
Conghui Wang
Yuwan Liu
Xinyi Wei
Mengxia Zheng
Jiaxin Gu
Xiaojing Chen
Yi Sun
Junfen Xu
Yan Lu
Weiguo Lu
author_sort Lingkai Gu
collection DOAJ
description Abstract Ovarian cancer is an aggressive gynecological tumor usually diagnosed with widespread metastases. Extracellular vesicles (EVs), though recognized as important mediators of tumor metastasis, have received limited attention into their specific functions via the mRNA profiling. Here it is reported elevated expression and selective enrichment of INAVA mRNA in both plasma‐ and tissue‐derived EVs from ovarian cancer patients, which is positively correlated with distant metastasis and poor prognosis. Functionally, INAVA mRNA, upon uptake and translation, activates normal ovarian fibroblasts (NOFs) and drives extensive peritoneum metastasis in the orthotopic xenograft mouse model. Mechanistically, INAVA competitively binds with high mobility group protein A2 (HMGA2) and consequently inhibit its interaction with vaccinia‐related kinase 1 (VRK1), leading to reduced HMGA2 phosphorylation on Ser105. Interestingly, this inhibitory phosphorylation stabilizes HMGA2 via blocking tripartite motif‐containing 21 (TRIM21) ‐mediated K48‐linked ubiquitylation, and ultimately enhances the transcription of STAT3 to activate NOFs. Lastly, a cell‐permeable peptide that disrupts the INAVA–HMGA2 interaction leads to attenuated NOF activation and provides a promising strategy for ovarian cancer therapy.
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spelling doaj-art-e482f754824d478ea5df6d0d998317702025-08-20T03:28:58ZengWileyAdvanced Science2198-38442025-07-011225n/an/a10.1002/advs.202500912The INAVA mRNA in Extracellular Vesicles Activates Normal Ovarian Fibroblasts by Phosphorylation–Ubiquitylation Crosstalk of HMGA2Lingkai Gu0Zhangjin Shen1Shizhen Shen2Conghui Wang3Yuwan Liu4Xinyi Wei5Mengxia Zheng6Jiaxin Gu7Xiaojing Chen8Yi Sun9Junfen Xu10Yan Lu11Weiguo Lu12Zhejiang Key Laboratory of Maternal and Infant Health Women's Hospital Zhejiang University School of Medicine Hangzhou 310006 ChinaDepartment of Obstetrics and Gynecology Sir Run Run Shaw Hospital School of Medicine Zhejiang University Hangzhou 310016 ChinaDepartment of Gynecologic Oncology Women's Hospital Zhejiang University School of Medicine Hangzhou 310006 ChinaDepartment of Gynecologic Oncology Women's Hospital Zhejiang University School of Medicine Hangzhou 310006 ChinaZhejiang Key Laboratory of Maternal and Infant Health Women's Hospital Zhejiang University School of Medicine Hangzhou 310006 ChinaDepartment of Gynecologic Oncology Women's Hospital Zhejiang University School of Medicine Hangzhou 310006 ChinaZhejiang Key Laboratory of Maternal and Infant Health Women's Hospital Zhejiang University School of Medicine Hangzhou 310006 ChinaZhejiang Key Laboratory of Maternal and Infant Health Women's Hospital Zhejiang University School of Medicine Hangzhou 310006 ChinaDepartment of Gynecologic Oncology Women's Hospital Zhejiang University School of Medicine Hangzhou 310006 ChinaInstitute of Translational Medicine Zhejiang University School of Medicine Hangzhou 310029 ChinaDepartment of Gynecologic Oncology Women's Hospital Zhejiang University School of Medicine Hangzhou 310006 ChinaDepartment of Gynecologic Oncology Women's Hospital Zhejiang University School of Medicine Hangzhou 310006 ChinaZhejiang Key Laboratory of Maternal and Infant Health Women's Hospital Zhejiang University School of Medicine Hangzhou 310006 ChinaAbstract Ovarian cancer is an aggressive gynecological tumor usually diagnosed with widespread metastases. Extracellular vesicles (EVs), though recognized as important mediators of tumor metastasis, have received limited attention into their specific functions via the mRNA profiling. Here it is reported elevated expression and selective enrichment of INAVA mRNA in both plasma‐ and tissue‐derived EVs from ovarian cancer patients, which is positively correlated with distant metastasis and poor prognosis. Functionally, INAVA mRNA, upon uptake and translation, activates normal ovarian fibroblasts (NOFs) and drives extensive peritoneum metastasis in the orthotopic xenograft mouse model. Mechanistically, INAVA competitively binds with high mobility group protein A2 (HMGA2) and consequently inhibit its interaction with vaccinia‐related kinase 1 (VRK1), leading to reduced HMGA2 phosphorylation on Ser105. Interestingly, this inhibitory phosphorylation stabilizes HMGA2 via blocking tripartite motif‐containing 21 (TRIM21) ‐mediated K48‐linked ubiquitylation, and ultimately enhances the transcription of STAT3 to activate NOFs. Lastly, a cell‐permeable peptide that disrupts the INAVA–HMGA2 interaction leads to attenuated NOF activation and provides a promising strategy for ovarian cancer therapy.https://doi.org/10.1002/advs.202500912extracellular vesiclesfibroblast activationinnate immunity activatorovarian cancerpost‐translational modifications interaction
spellingShingle Lingkai Gu
Zhangjin Shen
Shizhen Shen
Conghui Wang
Yuwan Liu
Xinyi Wei
Mengxia Zheng
Jiaxin Gu
Xiaojing Chen
Yi Sun
Junfen Xu
Yan Lu
Weiguo Lu
The INAVA mRNA in Extracellular Vesicles Activates Normal Ovarian Fibroblasts by Phosphorylation–Ubiquitylation Crosstalk of HMGA2
Advanced Science
extracellular vesicles
fibroblast activation
innate immunity activator
ovarian cancer
post‐translational modifications interaction
title The INAVA mRNA in Extracellular Vesicles Activates Normal Ovarian Fibroblasts by Phosphorylation–Ubiquitylation Crosstalk of HMGA2
title_full The INAVA mRNA in Extracellular Vesicles Activates Normal Ovarian Fibroblasts by Phosphorylation–Ubiquitylation Crosstalk of HMGA2
title_fullStr The INAVA mRNA in Extracellular Vesicles Activates Normal Ovarian Fibroblasts by Phosphorylation–Ubiquitylation Crosstalk of HMGA2
title_full_unstemmed The INAVA mRNA in Extracellular Vesicles Activates Normal Ovarian Fibroblasts by Phosphorylation–Ubiquitylation Crosstalk of HMGA2
title_short The INAVA mRNA in Extracellular Vesicles Activates Normal Ovarian Fibroblasts by Phosphorylation–Ubiquitylation Crosstalk of HMGA2
title_sort inava mrna in extracellular vesicles activates normal ovarian fibroblasts by phosphorylation ubiquitylation crosstalk of hmga2
topic extracellular vesicles
fibroblast activation
innate immunity activator
ovarian cancer
post‐translational modifications interaction
url https://doi.org/10.1002/advs.202500912
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