Prioritizing perturbation-responsive gene patterns using interpretable deep learning

Abstract Spatially resolved transcriptomics enables mapping of multiplexed gene expression within tissue contexts. While existing methods prioritize spatially variable genes within a single slice, few address identifying genes with differential spatial expression patterns (DSEPs) across multiple con...

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Bibliographic Details
Main Authors: Yan Cui, Zhiyuan Yuan
Format: Article
Language:English
Published: Nature Portfolio 2025-07-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-025-61476-9
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Summary:Abstract Spatially resolved transcriptomics enables mapping of multiplexed gene expression within tissue contexts. While existing methods prioritize spatially variable genes within a single slice, few address identifying genes with differential spatial expression patterns (DSEPs) across multiple conditions—an critical need for complex experimental designs. Challenges include modeling cross-slice spatial variation, scalability to large datasets, and disentangling inter-slice heterogeneity. We introduce DSEP gene prioritization as a new analytical task and present River, an interpretable deep learning framework that identifies genes exhibiting condition-relevant spatial changes. River features a two-branch predictive architecture and a post hoc attribution strategy to rank genes (or other features) by their contribution to condition differences. Its spatially-informed modeling ensures scalability to large spatial datasets, and we further decouple spatial and non-spatial components to enhance interpretability. We evaluate River on simulations and apply it to diverse biological contexts, including embryogenesis, diabetes-affected spermatogenesis, and lupus-associated splenic changes. In triple-negative breast cancer, River prioritizes survival-associated spatial patterns that generalize across patients. River is distribution-agnostic and compatible with diverse spatial data types, offering a flexible and scalable solution for analyzing tissue-wide expression dynamics across multiple biological conditions.
ISSN:2041-1723