RNA editing-induced structural and functional adaptations of NAD9 in Triticum aestivum under drought stress

IntroductionMitochondria are essential organelles in eukaryotic cells, producing ATP through the electron transport chain to supply energy for cellular activities. Beyond energy production, mitochondria play crucial roles in cellular signaling, stress responses, and the regulation of reactive oxygen...

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Main Authors: Nermin G. Mohamed, Ahmed M. Ramadan, Marwa Amer, Yasser Morsy, Rasha A. Mohamed, Osama A. M. Said, Afnan A. Alnufaei, Mona I M. Ibrahim, Sameh E. Hassanein, Hala F. Eissa
Format: Article
Language:English
Published: Frontiers Media S.A. 2024-11-01
Series:Frontiers in Plant Science
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Online Access:https://www.frontiersin.org/articles/10.3389/fpls.2024.1490288/full
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author Nermin G. Mohamed
Ahmed M. Ramadan
Ahmed M. Ramadan
Marwa Amer
Yasser Morsy
Yasser Morsy
Rasha A. Mohamed
Osama A. M. Said
Afnan A. Alnufaei
Mona I M. Ibrahim
Sameh E. Hassanein
Sameh E. Hassanein
Hala F. Eissa
Hala F. Eissa
author_facet Nermin G. Mohamed
Ahmed M. Ramadan
Ahmed M. Ramadan
Marwa Amer
Yasser Morsy
Yasser Morsy
Rasha A. Mohamed
Osama A. M. Said
Afnan A. Alnufaei
Mona I M. Ibrahim
Sameh E. Hassanein
Sameh E. Hassanein
Hala F. Eissa
Hala F. Eissa
author_sort Nermin G. Mohamed
collection DOAJ
description IntroductionMitochondria are essential organelles in eukaryotic cells, producing ATP through the electron transport chain to supply energy for cellular activities. Beyond energy production, mitochondria play crucial roles in cellular signaling, stress responses, and the regulation of reactive oxygen species. In plants, mitochondria are one of the keys to responding to environmental stresses which can significantly affect crop productivity, particularly in crops like wheat. RNA editing, a post-transcriptional RNA modification process in mitochondria, is linked to regulating these stress responses.MethodsThis study explores RNA editing patterns in the nad9 gene of wheat drought-tolerant (Giza168) and drought-sensitive (Gemmiza10) wheat cultivars under drought stress to understand plant adaptation mechanisms. RNA-seq data for these cultivars were analyzed using CLC Genomic Workbench to identify RNA editing sites in the nad9 gene, examining subsequent amino acid changes and predicting secondary structure modifications. These RNA editing sites were validated using qRT-PCR on drought-treated seedlings at 0, 2, and 12 hours post-treatment. Protein models were generated using AlphaFold, with functional predictions and structure verification conducted using various bioinformatics tools to investigate the effect of RNA editing on protein level.ResultsThe results showed significant RNA editing events, especially C-to-T conversions, in the nad9 gene across different drought exposure times. Giza168 had 22 editing sites, while Gemmiza10 had 19, with several showing significant differences between control and stress conditions. RNA editing influenced the NAD9 protein's secondary structure, particularly beta sheets, and 3D modeling highlighted the structural impacts of these edits. The N-terminal region of NAD9 contained important regulatory motifs, suggesting a complex regulatory environment.DiscussionThis study reveals key editing sites that differ between drought-tolerant and sensitive wheat cultivars, impacting NAD9 protein structures and highlighting the role of RNA editing in enhancing drought resilience. Additionally, the study suggests potential regulatory mechanisms, including phosphorylation and ubiquitination that influence mitochondrial stability and function.
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spelling doaj-art-e3f6f7b094014bcdbe843d8c02c08d702025-08-20T02:14:49ZengFrontiers Media S.A.Frontiers in Plant Science1664-462X2024-11-011510.3389/fpls.2024.14902881490288RNA editing-induced structural and functional adaptations of NAD9 in Triticum aestivum under drought stressNermin G. Mohamed0Ahmed M. Ramadan1Ahmed M. Ramadan2Marwa Amer3Yasser Morsy4Yasser Morsy5Rasha A. Mohamed6Osama A. M. Said7Afnan A. Alnufaei8Mona I M. Ibrahim9Sameh E. Hassanein10Sameh E. Hassanein11Hala F. Eissa12Hala F. Eissa13College of Biotechnology, Misr University for Science and Technology (MUST), Giza, EgyptBiological Science Department, Faculty of Science, King Abdulaziz University, Jeddah, Saudi ArabiaPrincess Najla bint Saud Al-Saud Center for Excellence Research in Biotechnology, King Abdulaziz University, Jeddah, Saudi ArabiaCollege of Biotechnology, Misr University for Science and Technology (MUST), Giza, EgyptDepartment of Gastroenterology and Hepatology, University Hospital Zurich, University of Zurich, Zurich, SwitzerlandAgricultural Genetic Engineering Research Institute (AGERI), Agricultural Research Center (ARC), Giza, EgyptAgricultural Genetic Engineering Research Institute (AGERI), Agricultural Research Center (ARC), Giza, EgyptCollege of Biotechnology, Misr University for Science and Technology (MUST), Giza, EgyptDepartment of Biology, Faculty of Science, University of Bisha, Bisha, Saudi ArabiaCollege of Biotechnology, Misr University for Science and Technology (MUST), Giza, EgyptAgricultural Genetic Engineering Research Institute (AGERI), Agricultural Research Center (ARC), Giza, EgyptBioinformatics Program, School of Biotechnology, Nile University, Giza, EgyptCollege of Biotechnology, Misr University for Science and Technology (MUST), Giza, EgyptAgricultural Genetic Engineering Research Institute (AGERI), Agricultural Research Center (ARC), Giza, EgyptIntroductionMitochondria are essential organelles in eukaryotic cells, producing ATP through the electron transport chain to supply energy for cellular activities. Beyond energy production, mitochondria play crucial roles in cellular signaling, stress responses, and the regulation of reactive oxygen species. In plants, mitochondria are one of the keys to responding to environmental stresses which can significantly affect crop productivity, particularly in crops like wheat. RNA editing, a post-transcriptional RNA modification process in mitochondria, is linked to regulating these stress responses.MethodsThis study explores RNA editing patterns in the nad9 gene of wheat drought-tolerant (Giza168) and drought-sensitive (Gemmiza10) wheat cultivars under drought stress to understand plant adaptation mechanisms. RNA-seq data for these cultivars were analyzed using CLC Genomic Workbench to identify RNA editing sites in the nad9 gene, examining subsequent amino acid changes and predicting secondary structure modifications. These RNA editing sites were validated using qRT-PCR on drought-treated seedlings at 0, 2, and 12 hours post-treatment. Protein models were generated using AlphaFold, with functional predictions and structure verification conducted using various bioinformatics tools to investigate the effect of RNA editing on protein level.ResultsThe results showed significant RNA editing events, especially C-to-T conversions, in the nad9 gene across different drought exposure times. Giza168 had 22 editing sites, while Gemmiza10 had 19, with several showing significant differences between control and stress conditions. RNA editing influenced the NAD9 protein's secondary structure, particularly beta sheets, and 3D modeling highlighted the structural impacts of these edits. The N-terminal region of NAD9 contained important regulatory motifs, suggesting a complex regulatory environment.DiscussionThis study reveals key editing sites that differ between drought-tolerant and sensitive wheat cultivars, impacting NAD9 protein structures and highlighting the role of RNA editing in enhancing drought resilience. Additionally, the study suggests potential regulatory mechanisms, including phosphorylation and ubiquitination that influence mitochondrial stability and function.https://www.frontiersin.org/articles/10.3389/fpls.2024.1490288/fullRNA editingnad9 genedrought stressTriticum aestivumpost-translational modification
spellingShingle Nermin G. Mohamed
Ahmed M. Ramadan
Ahmed M. Ramadan
Marwa Amer
Yasser Morsy
Yasser Morsy
Rasha A. Mohamed
Osama A. M. Said
Afnan A. Alnufaei
Mona I M. Ibrahim
Sameh E. Hassanein
Sameh E. Hassanein
Hala F. Eissa
Hala F. Eissa
RNA editing-induced structural and functional adaptations of NAD9 in Triticum aestivum under drought stress
Frontiers in Plant Science
RNA editing
nad9 gene
drought stress
Triticum aestivum
post-translational modification
title RNA editing-induced structural and functional adaptations of NAD9 in Triticum aestivum under drought stress
title_full RNA editing-induced structural and functional adaptations of NAD9 in Triticum aestivum under drought stress
title_fullStr RNA editing-induced structural and functional adaptations of NAD9 in Triticum aestivum under drought stress
title_full_unstemmed RNA editing-induced structural and functional adaptations of NAD9 in Triticum aestivum under drought stress
title_short RNA editing-induced structural and functional adaptations of NAD9 in Triticum aestivum under drought stress
title_sort rna editing induced structural and functional adaptations of nad9 in triticum aestivum under drought stress
topic RNA editing
nad9 gene
drought stress
Triticum aestivum
post-translational modification
url https://www.frontiersin.org/articles/10.3389/fpls.2024.1490288/full
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