Phosphatidylinositides regulate the cell plate morphology transition during cytokinesis in Arabidopsis

Abstract In plants, the developing cell plate which is characterized by a series of anionic lipids, undergoes dramatic morphological change for successful cytokinesis. However, the mechanisms underlying these alterations, and the roles of anionic lipids such as phosphatidylinositol-4-phosphate (PI4P...

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Main Authors: Yu Luo, Yu-Fang Tian, Hui-Ru Liu, Wei-Cai Yang
Format: Article
Language:English
Published: Nature Portfolio 2025-07-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-025-62067-4
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author Yu Luo
Yu-Fang Tian
Hui-Ru Liu
Wei-Cai Yang
author_facet Yu Luo
Yu-Fang Tian
Hui-Ru Liu
Wei-Cai Yang
author_sort Yu Luo
collection DOAJ
description Abstract In plants, the developing cell plate which is characterized by a series of anionic lipids, undergoes dramatic morphological change for successful cytokinesis. However, the mechanisms underlying these alterations, and the roles of anionic lipids such as phosphatidylinositol-4-phosphate (PI4P), phosphatidylinositol-4,5-bisphosphate (PI(4,5)P2), and phosphatidylserine (PS) during cell division remain poorly understood. Here we present that changes in anionic lipid composition have a profound effect on cell plate development: deprivation of phosphatidylinositides (PIPs) leads to incomplete cytokinesis through distorted cell-plate architecture. Our data demonstrate that PI4P shapes cell plate membrane morphology through flippase-regulated PS flipping inhibition, while PI(4,5)P2 functions in the recruitment of dynamin-related protein 1A (DRP1A) and the constriction region formation; depletion of PIPs causes cell plate tubulation and flattening failure. We propose a model in which PI4P regulates the level and distribution of PS, while PI(4,5)P2 mediates the localization of DRP1A; together, they coordinate cell plate morphology to ensure successful cytokinesis in plant cells.
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spelling doaj-art-e32ee5dc84414a16b2833294defdcdd12025-08-20T03:43:14ZengNature PortfolioNature Communications2041-17232025-07-0116111610.1038/s41467-025-62067-4Phosphatidylinositides regulate the cell plate morphology transition during cytokinesis in ArabidopsisYu Luo0Yu-Fang Tian1Hui-Ru Liu2Wei-Cai Yang3State Key Laboratory of Molecular and Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of SciencesState Key Laboratory of Molecular and Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of SciencesState Key Laboratory of Molecular and Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of SciencesState Key Laboratory of Molecular and Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of SciencesAbstract In plants, the developing cell plate which is characterized by a series of anionic lipids, undergoes dramatic morphological change for successful cytokinesis. However, the mechanisms underlying these alterations, and the roles of anionic lipids such as phosphatidylinositol-4-phosphate (PI4P), phosphatidylinositol-4,5-bisphosphate (PI(4,5)P2), and phosphatidylserine (PS) during cell division remain poorly understood. Here we present that changes in anionic lipid composition have a profound effect on cell plate development: deprivation of phosphatidylinositides (PIPs) leads to incomplete cytokinesis through distorted cell-plate architecture. Our data demonstrate that PI4P shapes cell plate membrane morphology through flippase-regulated PS flipping inhibition, while PI(4,5)P2 functions in the recruitment of dynamin-related protein 1A (DRP1A) and the constriction region formation; depletion of PIPs causes cell plate tubulation and flattening failure. We propose a model in which PI4P regulates the level and distribution of PS, while PI(4,5)P2 mediates the localization of DRP1A; together, they coordinate cell plate morphology to ensure successful cytokinesis in plant cells.https://doi.org/10.1038/s41467-025-62067-4
spellingShingle Yu Luo
Yu-Fang Tian
Hui-Ru Liu
Wei-Cai Yang
Phosphatidylinositides regulate the cell plate morphology transition during cytokinesis in Arabidopsis
Nature Communications
title Phosphatidylinositides regulate the cell plate morphology transition during cytokinesis in Arabidopsis
title_full Phosphatidylinositides regulate the cell plate morphology transition during cytokinesis in Arabidopsis
title_fullStr Phosphatidylinositides regulate the cell plate morphology transition during cytokinesis in Arabidopsis
title_full_unstemmed Phosphatidylinositides regulate the cell plate morphology transition during cytokinesis in Arabidopsis
title_short Phosphatidylinositides regulate the cell plate morphology transition during cytokinesis in Arabidopsis
title_sort phosphatidylinositides regulate the cell plate morphology transition during cytokinesis in arabidopsis
url https://doi.org/10.1038/s41467-025-62067-4
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AT huiruliu phosphatidylinositidesregulatethecellplatemorphologytransitionduringcytokinesisinarabidopsis
AT weicaiyang phosphatidylinositidesregulatethecellplatemorphologytransitionduringcytokinesisinarabidopsis