Foxk1 and Foxk2 promote cardiomyocyte proliferation and heart regeneration

Abstract Promoting endogenous cardiomyocyte proliferation is a promising strategy for cardiac repair. Identifying key factors that regulate cardiomyocyte proliferation can advance the development of novel therapies for heart regeneration. Here, we identify Foxk1 and Foxk2 as key regulators of cardio...

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Main Authors: Dongcheng Cai, Chungeng Liu, Haotong Li, Chiyin Wang, Lina Bai, Jie Feng, Miaoqing Hu, Hao Wang, Shen Song, Yifan Xie, Ziwei Chen, Jiajun Zhong, Hong Lian, Zhiwei Yang, Yuhui Zhang, Yu Nie
Format: Article
Language:English
Published: Nature Portfolio 2025-03-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-025-57996-z
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author Dongcheng Cai
Chungeng Liu
Haotong Li
Chiyin Wang
Lina Bai
Jie Feng
Miaoqing Hu
Hao Wang
Shen Song
Yifan Xie
Ziwei Chen
Jiajun Zhong
Hong Lian
Zhiwei Yang
Yuhui Zhang
Yu Nie
author_facet Dongcheng Cai
Chungeng Liu
Haotong Li
Chiyin Wang
Lina Bai
Jie Feng
Miaoqing Hu
Hao Wang
Shen Song
Yifan Xie
Ziwei Chen
Jiajun Zhong
Hong Lian
Zhiwei Yang
Yuhui Zhang
Yu Nie
author_sort Dongcheng Cai
collection DOAJ
description Abstract Promoting endogenous cardiomyocyte proliferation is a promising strategy for cardiac repair. Identifying key factors that regulate cardiomyocyte proliferation can advance the development of novel therapies for heart regeneration. Here, we identify Foxk1 and Foxk2 as key regulators of cardiomyocyte proliferation, whose expression declines during postnatal heart development. Cardiomyocyte-specific knockout of Foxk1 or Foxk2 impairs neonatal heart regeneration after myocardial infarction (MI) injury. AAV9-mediated Foxk1 or Foxk2 overexpression extends the postnatal cardiomyocyte proliferative window and enhances cardiac repair in adult mice after MI. Mechanistically, Foxk1 and Foxk2 drive cardiomyocyte cell cycle progression by directly activating CCNB1 and CDK1 expression, forming the CCNB1/CDK1 complex that facilitates G2/M transition. Moreover, Foxk1 and Foxk2 promote cardiomyocyte proliferation by upregulating HIF1α expression, which enhances glycolysis and the pentose phosphate pathway (PPP), which further favors cardiomyocyte proliferation. These findings establish Foxk1 and Foxk2 as promising therapeutic targets for cardiac injury.
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institution Kabale University
issn 2041-1723
language English
publishDate 2025-03-01
publisher Nature Portfolio
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series Nature Communications
spelling doaj-art-e2e41d7b403c4967a5967b1d297985b52025-08-20T03:40:45ZengNature PortfolioNature Communications2041-17232025-03-0116111710.1038/s41467-025-57996-zFoxk1 and Foxk2 promote cardiomyocyte proliferation and heart regenerationDongcheng Cai0Chungeng Liu1Haotong Li2Chiyin Wang3Lina Bai4Jie Feng5Miaoqing Hu6Hao Wang7Shen Song8Yifan Xie9Ziwei Chen10Jiajun Zhong11Hong Lian12Zhiwei Yang13Yuhui Zhang14Yu Nie15State Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeNational Health Commission Key Laboratory of Human Disease Comparative Medicine, Institute of Laboratory Animal Sciences, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeState Key Laboratory of Cardiovascular Disease, Fuwai Hospital, National Center for Cardiovascular Disease, Chinese Academy of Medical Sciences and Peking Union Medical CollegeAbstract Promoting endogenous cardiomyocyte proliferation is a promising strategy for cardiac repair. Identifying key factors that regulate cardiomyocyte proliferation can advance the development of novel therapies for heart regeneration. Here, we identify Foxk1 and Foxk2 as key regulators of cardiomyocyte proliferation, whose expression declines during postnatal heart development. Cardiomyocyte-specific knockout of Foxk1 or Foxk2 impairs neonatal heart regeneration after myocardial infarction (MI) injury. AAV9-mediated Foxk1 or Foxk2 overexpression extends the postnatal cardiomyocyte proliferative window and enhances cardiac repair in adult mice after MI. Mechanistically, Foxk1 and Foxk2 drive cardiomyocyte cell cycle progression by directly activating CCNB1 and CDK1 expression, forming the CCNB1/CDK1 complex that facilitates G2/M transition. Moreover, Foxk1 and Foxk2 promote cardiomyocyte proliferation by upregulating HIF1α expression, which enhances glycolysis and the pentose phosphate pathway (PPP), which further favors cardiomyocyte proliferation. These findings establish Foxk1 and Foxk2 as promising therapeutic targets for cardiac injury.https://doi.org/10.1038/s41467-025-57996-z
spellingShingle Dongcheng Cai
Chungeng Liu
Haotong Li
Chiyin Wang
Lina Bai
Jie Feng
Miaoqing Hu
Hao Wang
Shen Song
Yifan Xie
Ziwei Chen
Jiajun Zhong
Hong Lian
Zhiwei Yang
Yuhui Zhang
Yu Nie
Foxk1 and Foxk2 promote cardiomyocyte proliferation and heart regeneration
Nature Communications
title Foxk1 and Foxk2 promote cardiomyocyte proliferation and heart regeneration
title_full Foxk1 and Foxk2 promote cardiomyocyte proliferation and heart regeneration
title_fullStr Foxk1 and Foxk2 promote cardiomyocyte proliferation and heart regeneration
title_full_unstemmed Foxk1 and Foxk2 promote cardiomyocyte proliferation and heart regeneration
title_short Foxk1 and Foxk2 promote cardiomyocyte proliferation and heart regeneration
title_sort foxk1 and foxk2 promote cardiomyocyte proliferation and heart regeneration
url https://doi.org/10.1038/s41467-025-57996-z
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