Nuclear p62/SQSTM1 facilitates ubiquitin-independent proteasomal degradation of BMAL1.

Brain and muscle arnt-like protein 1(BMAL1) is a critical regulator of circadian rhythm. Although transcriptional regulation of BMAL1 has been extensively studied, the mechanisms governing the stability of BMAL1 at the protein level remain unclear. p62/SQSTM1 is a crucial factor in proteostasis regu...

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Main Authors: Chenliang Zhang, Quanyou Wu, Huan Zhang, Ruichen Liu, Liping Li
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2025-07-01
Series:PLoS Genetics
Online Access:https://doi.org/10.1371/journal.pgen.1011794
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author Chenliang Zhang
Quanyou Wu
Huan Zhang
Ruichen Liu
Liping Li
author_facet Chenliang Zhang
Quanyou Wu
Huan Zhang
Ruichen Liu
Liping Li
author_sort Chenliang Zhang
collection DOAJ
description Brain and muscle arnt-like protein 1(BMAL1) is a critical regulator of circadian rhythm. Although transcriptional regulation of BMAL1 has been extensively studied, the mechanisms governing the stability of BMAL1 at the protein level remain unclear. p62/SQSTM1 is a crucial factor in proteostasis regulation and is involved in both autophagy and the ubiquitin-proteasome system. We demonstrated that p62 promotes proteasomal degradation of BMAL1 within the nucleus, independent of ubiquitination. Additional molecular analyses indicated that p62 functions as a receptor for the 20S proteasome, facilitating the recruitment of BMAL1 to the 20S proteasome for degradation. This mechanism is independent of recently identified p62-driven nuclear biomolecular condensates. We also revealed that remodeling the nuclear accumulation of p62 may represent a potential strategy for targeting BMAL1 to suppress tumor cell growth. In conclusion, our findings revealed a novel mechanism by which nuclear p62 regulates BMAL1 proteostasis.
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institution Kabale University
issn 1553-7390
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language English
publishDate 2025-07-01
publisher Public Library of Science (PLoS)
record_format Article
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spelling doaj-art-e1fcb0f66bdb4df69705c71d93ad29782025-08-20T03:51:09ZengPublic Library of Science (PLoS)PLoS Genetics1553-73901553-74042025-07-01217e101179410.1371/journal.pgen.1011794Nuclear p62/SQSTM1 facilitates ubiquitin-independent proteasomal degradation of BMAL1.Chenliang ZhangQuanyou WuHuan ZhangRuichen LiuLiping LiBrain and muscle arnt-like protein 1(BMAL1) is a critical regulator of circadian rhythm. Although transcriptional regulation of BMAL1 has been extensively studied, the mechanisms governing the stability of BMAL1 at the protein level remain unclear. p62/SQSTM1 is a crucial factor in proteostasis regulation and is involved in both autophagy and the ubiquitin-proteasome system. We demonstrated that p62 promotes proteasomal degradation of BMAL1 within the nucleus, independent of ubiquitination. Additional molecular analyses indicated that p62 functions as a receptor for the 20S proteasome, facilitating the recruitment of BMAL1 to the 20S proteasome for degradation. This mechanism is independent of recently identified p62-driven nuclear biomolecular condensates. We also revealed that remodeling the nuclear accumulation of p62 may represent a potential strategy for targeting BMAL1 to suppress tumor cell growth. In conclusion, our findings revealed a novel mechanism by which nuclear p62 regulates BMAL1 proteostasis.https://doi.org/10.1371/journal.pgen.1011794
spellingShingle Chenliang Zhang
Quanyou Wu
Huan Zhang
Ruichen Liu
Liping Li
Nuclear p62/SQSTM1 facilitates ubiquitin-independent proteasomal degradation of BMAL1.
PLoS Genetics
title Nuclear p62/SQSTM1 facilitates ubiquitin-independent proteasomal degradation of BMAL1.
title_full Nuclear p62/SQSTM1 facilitates ubiquitin-independent proteasomal degradation of BMAL1.
title_fullStr Nuclear p62/SQSTM1 facilitates ubiquitin-independent proteasomal degradation of BMAL1.
title_full_unstemmed Nuclear p62/SQSTM1 facilitates ubiquitin-independent proteasomal degradation of BMAL1.
title_short Nuclear p62/SQSTM1 facilitates ubiquitin-independent proteasomal degradation of BMAL1.
title_sort nuclear p62 sqstm1 facilitates ubiquitin independent proteasomal degradation of bmal1
url https://doi.org/10.1371/journal.pgen.1011794
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AT huanzhang nuclearp62sqstm1facilitatesubiquitinindependentproteasomaldegradationofbmal1
AT ruichenliu nuclearp62sqstm1facilitatesubiquitinindependentproteasomaldegradationofbmal1
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