A plant Bro1 domain protein BRAF regulates multivesicular body biogenesis and membrane protein homeostasis

Abstract Plant development, defense, and many physiological processes rely on the endosomal sorting complex required for transport (ESCRT) machinery to control the homeostasis of membrane proteins by selective vacuolar degradation. Although ESCRT core components are conserved among higher eukaryotes...

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Main Authors: Jinbo Shen, Qiong Zhao, Xiangfeng Wang, Caiji Gao, Ying Zhu, Yonglun Zeng, Liwen Jiang
Format: Article
Language:English
Published: Nature Portfolio 2018-09-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-018-05913-y
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author Jinbo Shen
Qiong Zhao
Xiangfeng Wang
Caiji Gao
Ying Zhu
Yonglun Zeng
Liwen Jiang
author_facet Jinbo Shen
Qiong Zhao
Xiangfeng Wang
Caiji Gao
Ying Zhu
Yonglun Zeng
Liwen Jiang
author_sort Jinbo Shen
collection DOAJ
description Abstract Plant development, defense, and many physiological processes rely on the endosomal sorting complex required for transport (ESCRT) machinery to control the homeostasis of membrane proteins by selective vacuolar degradation. Although ESCRT core components are conserved among higher eukaryotes, the regulators that control the function of the ESCRT machinery remain elusive. We recently identified a plant-specific ESCRT component, FREE1, that is essential for multivesicular body/prevacuolar compartment (MVB/PVC) biogenesis and vacuolar sorting of membrane proteins. Here we identify a plant-specific Bro1-domain protein BRAF, which regulates FREE1 recruitment to the MVB/PVC membrane by competitively binding to the ESCRT-I component Vps23. Altogether, we have successfully identified a role for BRAF, whose function as a unique evolutionary ESCRT regulator in orchestrating intraluminal vesicle formation in MVB/PVCs and the sorting of membrane proteins for degradation in plants makes it an important regulatory mechanism underlying the ESCRT machinery in higher eukaryotes.
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issn 2041-1723
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spelling doaj-art-e07d09322bbf42ac97b3fb0813458d2b2025-08-20T02:34:04ZengNature PortfolioNature Communications2041-17232018-09-019111510.1038/s41467-018-05913-yA plant Bro1 domain protein BRAF regulates multivesicular body biogenesis and membrane protein homeostasisJinbo Shen0Qiong Zhao1Xiangfeng Wang2Caiji Gao3Ying Zhu4Yonglun Zeng5Liwen Jiang6Centre for Cell & Developmental Biology, State Key Laboratory of Agrobiotechnology, School of Life Sciences, The Chinese University of Hong KongCentre for Cell & Developmental Biology, State Key Laboratory of Agrobiotechnology, School of Life Sciences, The Chinese University of Hong KongCentre for Cell & Developmental Biology, State Key Laboratory of Agrobiotechnology, School of Life Sciences, The Chinese University of Hong KongCentre for Cell & Developmental Biology, State Key Laboratory of Agrobiotechnology, School of Life Sciences, The Chinese University of Hong KongCentre for Cell & Developmental Biology, State Key Laboratory of Agrobiotechnology, School of Life Sciences, The Chinese University of Hong KongCentre for Cell & Developmental Biology, State Key Laboratory of Agrobiotechnology, School of Life Sciences, The Chinese University of Hong KongCentre for Cell & Developmental Biology, State Key Laboratory of Agrobiotechnology, School of Life Sciences, The Chinese University of Hong KongAbstract Plant development, defense, and many physiological processes rely on the endosomal sorting complex required for transport (ESCRT) machinery to control the homeostasis of membrane proteins by selective vacuolar degradation. Although ESCRT core components are conserved among higher eukaryotes, the regulators that control the function of the ESCRT machinery remain elusive. We recently identified a plant-specific ESCRT component, FREE1, that is essential for multivesicular body/prevacuolar compartment (MVB/PVC) biogenesis and vacuolar sorting of membrane proteins. Here we identify a plant-specific Bro1-domain protein BRAF, which regulates FREE1 recruitment to the MVB/PVC membrane by competitively binding to the ESCRT-I component Vps23. Altogether, we have successfully identified a role for BRAF, whose function as a unique evolutionary ESCRT regulator in orchestrating intraluminal vesicle formation in MVB/PVCs and the sorting of membrane proteins for degradation in plants makes it an important regulatory mechanism underlying the ESCRT machinery in higher eukaryotes.https://doi.org/10.1038/s41467-018-05913-y
spellingShingle Jinbo Shen
Qiong Zhao
Xiangfeng Wang
Caiji Gao
Ying Zhu
Yonglun Zeng
Liwen Jiang
A plant Bro1 domain protein BRAF regulates multivesicular body biogenesis and membrane protein homeostasis
Nature Communications
title A plant Bro1 domain protein BRAF regulates multivesicular body biogenesis and membrane protein homeostasis
title_full A plant Bro1 domain protein BRAF regulates multivesicular body biogenesis and membrane protein homeostasis
title_fullStr A plant Bro1 domain protein BRAF regulates multivesicular body biogenesis and membrane protein homeostasis
title_full_unstemmed A plant Bro1 domain protein BRAF regulates multivesicular body biogenesis and membrane protein homeostasis
title_short A plant Bro1 domain protein BRAF regulates multivesicular body biogenesis and membrane protein homeostasis
title_sort plant bro1 domain protein braf regulates multivesicular body biogenesis and membrane protein homeostasis
url https://doi.org/10.1038/s41467-018-05913-y
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