Transcriptomic analysis reveals the mechanism underlying salinity-induced morphological changes in Skeletonema subsalsum

Diatom cell walls are diverse and unique, providing the basis for species identification and supporting the ecological and economic value of diatoms. However, these important structures sometimes change in response to environmental fluctuations, especially under salt adaptation. Although studies hav...

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Main Authors: Jingwen Hu, Ya Zheng, Shuang Yang, Lin Yang, Qingmin You, Quanxi Wang
Format: Article
Language:English
Published: Frontiers Media S.A. 2024-10-01
Series:Frontiers in Microbiology
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Online Access:https://www.frontiersin.org/articles/10.3389/fmicb.2024.1476738/full
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author Jingwen Hu
Ya Zheng
Shuang Yang
Lin Yang
Lin Yang
Qingmin You
Quanxi Wang
author_facet Jingwen Hu
Ya Zheng
Shuang Yang
Lin Yang
Lin Yang
Qingmin You
Quanxi Wang
author_sort Jingwen Hu
collection DOAJ
description Diatom cell walls are diverse and unique, providing the basis for species identification and supporting the ecological and economic value of diatoms. However, these important structures sometimes change in response to environmental fluctuations, especially under salt adaptation. Although studies have shown that salinity induces morphological plasticity changes in diatom cell walls, most research has focused on physiological responses rather than molecular mechanisms. In this study, Skeletonema subsalsum was cultured under four salinity conditions (0, 3, 6, 12). Through morphological and physiological methods, we found that salinity increased the cell diameter, protrusion lengths, distance between adjacent cells (DBCs), and nanopore size, while reducing cell height and silicification degree. To further investigate the mechanism underlying morphological changes in S. subsalsum, complementary transcriptome analysis was performed. In total, 20,138 differentially expressed genes (DEGs) were identified among the four treatments. Among them, 231 DEGs were screened and found to be closely associated with morphological changes, of which 107 were downregulated and 124 were upregulated. The findings demonstrated that elevated salinity inhibited silicon transport and deposition via downregulating the expression of DEGs involved in functions including chitin metabolism, putrescine metabolism, and vesicle transport, resulting in reduced silicon content and cell height. Increased salinity promoted the expression of DEGs related to microtubules (MTs), actin, and ubiquitin, which synchronously induced morphological changes. These findings provide a more comprehensive understanding of the salt tolerance of algae and a foundation for future studies on cell wall morphogenesis.
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spelling doaj-art-ddd38c6c0ed94bd2911cd938e4eef9df2025-08-20T02:12:10ZengFrontiers Media S.A.Frontiers in Microbiology1664-302X2024-10-011510.3389/fmicb.2024.14767381476738Transcriptomic analysis reveals the mechanism underlying salinity-induced morphological changes in Skeletonema subsalsumJingwen Hu0Ya Zheng1Shuang Yang2Lin Yang3Lin Yang4Qingmin You5Quanxi Wang6Laboratory of Algae and Environment, College of Life Sciences, Shanghai Normal University, Shanghai, ChinaLaboratory of Algae and Environment, College of Life Sciences, Shanghai Normal University, Shanghai, ChinaLaboratory of Algae and Environment, College of Life Sciences, Shanghai Normal University, Shanghai, ChinaLaboratory of Algae and Environment, College of Life Sciences, Shanghai Normal University, Shanghai, ChinaLaboratory of Environmental Ecology and Engineering, College of Life Sciences, Hengshui University, Hengshui, ChinaLaboratory of Algae and Environment, College of Life Sciences, Shanghai Normal University, Shanghai, ChinaLaboratory of Algae and Environment, College of Life Sciences, Shanghai Normal University, Shanghai, ChinaDiatom cell walls are diverse and unique, providing the basis for species identification and supporting the ecological and economic value of diatoms. However, these important structures sometimes change in response to environmental fluctuations, especially under salt adaptation. Although studies have shown that salinity induces morphological plasticity changes in diatom cell walls, most research has focused on physiological responses rather than molecular mechanisms. In this study, Skeletonema subsalsum was cultured under four salinity conditions (0, 3, 6, 12). Through morphological and physiological methods, we found that salinity increased the cell diameter, protrusion lengths, distance between adjacent cells (DBCs), and nanopore size, while reducing cell height and silicification degree. To further investigate the mechanism underlying morphological changes in S. subsalsum, complementary transcriptome analysis was performed. In total, 20,138 differentially expressed genes (DEGs) were identified among the four treatments. Among them, 231 DEGs were screened and found to be closely associated with morphological changes, of which 107 were downregulated and 124 were upregulated. The findings demonstrated that elevated salinity inhibited silicon transport and deposition via downregulating the expression of DEGs involved in functions including chitin metabolism, putrescine metabolism, and vesicle transport, resulting in reduced silicon content and cell height. Increased salinity promoted the expression of DEGs related to microtubules (MTs), actin, and ubiquitin, which synchronously induced morphological changes. These findings provide a more comprehensive understanding of the salt tolerance of algae and a foundation for future studies on cell wall morphogenesis.https://www.frontiersin.org/articles/10.3389/fmicb.2024.1476738/fulldiatomssalinitymorphological changescell wall morphogenesismolecular mechanism
spellingShingle Jingwen Hu
Ya Zheng
Shuang Yang
Lin Yang
Lin Yang
Qingmin You
Quanxi Wang
Transcriptomic analysis reveals the mechanism underlying salinity-induced morphological changes in Skeletonema subsalsum
Frontiers in Microbiology
diatoms
salinity
morphological changes
cell wall morphogenesis
molecular mechanism
title Transcriptomic analysis reveals the mechanism underlying salinity-induced morphological changes in Skeletonema subsalsum
title_full Transcriptomic analysis reveals the mechanism underlying salinity-induced morphological changes in Skeletonema subsalsum
title_fullStr Transcriptomic analysis reveals the mechanism underlying salinity-induced morphological changes in Skeletonema subsalsum
title_full_unstemmed Transcriptomic analysis reveals the mechanism underlying salinity-induced morphological changes in Skeletonema subsalsum
title_short Transcriptomic analysis reveals the mechanism underlying salinity-induced morphological changes in Skeletonema subsalsum
title_sort transcriptomic analysis reveals the mechanism underlying salinity induced morphological changes in skeletonema subsalsum
topic diatoms
salinity
morphological changes
cell wall morphogenesis
molecular mechanism
url https://www.frontiersin.org/articles/10.3389/fmicb.2024.1476738/full
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