Structural and Biochemical Characterization of a Widespread Enterobacterial Peroxidase Encapsulin

Abstract Encapsulins are self‐assembling protein compartments found in prokaryotes and specifically encapsulate dedicated cargo enzymes. The most abundant encapsulin cargo class are Dye‐decolorizing Peroxidases (DyPs). It has been previously suggested that DyP encapsulins are involved in oxidative s...

Full description

Saved in:
Bibliographic Details
Main Authors: Natalia C. Ubilla‐Rodriguez, Michael P. Andreas, Tobias W. Giessen
Format: Article
Language:English
Published: Wiley 2025-06-01
Series:Advanced Science
Subjects:
Online Access:https://doi.org/10.1002/advs.202415827
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1850131356203352064
author Natalia C. Ubilla‐Rodriguez
Michael P. Andreas
Tobias W. Giessen
author_facet Natalia C. Ubilla‐Rodriguez
Michael P. Andreas
Tobias W. Giessen
author_sort Natalia C. Ubilla‐Rodriguez
collection DOAJ
description Abstract Encapsulins are self‐assembling protein compartments found in prokaryotes and specifically encapsulate dedicated cargo enzymes. The most abundant encapsulin cargo class are Dye‐decolorizing Peroxidases (DyPs). It has been previously suggested that DyP encapsulins are involved in oxidative stress resistance and bacterial pathogenicity due to DyPs’ inherent ability to reduce and detoxify hydrogen peroxide while oxidizing a broad range of organic co‐substrates. Here, we report the structural and biochemical analysis of a DyP encapsulin widely found across enterobacteria. Using bioinformatic approaches, we show that this DyP encapsulin is encoded by a conserved transposon‐associated operon, enriched in enterobacterial pathogens. Through low pH and peroxide exposure experiments, we highlight the stability of this DyP encapsulin under harsh conditions and show that DyP catalytic activity is highest at low pH. We determine the structure of the DyP‐loaded shell and free DyP via cryo‐electron microscopy, revealing the structural basis for DyP cargo loading and peroxide preference. This work lays the foundation to further explore the substrate range and physiological functions of enterobacterial DyP encapsulins.
format Article
id doaj-art-dda44a84d83644f698f9a6a42b99e324
institution OA Journals
issn 2198-3844
language English
publishDate 2025-06-01
publisher Wiley
record_format Article
series Advanced Science
spelling doaj-art-dda44a84d83644f698f9a6a42b99e3242025-08-20T02:32:26ZengWileyAdvanced Science2198-38442025-06-011221n/an/a10.1002/advs.202415827Structural and Biochemical Characterization of a Widespread Enterobacterial Peroxidase EncapsulinNatalia C. Ubilla‐Rodriguez0Michael P. Andreas1Tobias W. Giessen2Department of Biological Chemistry University of Michigan Medical School Ann Arbor MI 48109 USADepartment of Biological Chemistry University of Michigan Medical School Ann Arbor MI 48109 USADepartment of Biological Chemistry University of Michigan Medical School Ann Arbor MI 48109 USAAbstract Encapsulins are self‐assembling protein compartments found in prokaryotes and specifically encapsulate dedicated cargo enzymes. The most abundant encapsulin cargo class are Dye‐decolorizing Peroxidases (DyPs). It has been previously suggested that DyP encapsulins are involved in oxidative stress resistance and bacterial pathogenicity due to DyPs’ inherent ability to reduce and detoxify hydrogen peroxide while oxidizing a broad range of organic co‐substrates. Here, we report the structural and biochemical analysis of a DyP encapsulin widely found across enterobacteria. Using bioinformatic approaches, we show that this DyP encapsulin is encoded by a conserved transposon‐associated operon, enriched in enterobacterial pathogens. Through low pH and peroxide exposure experiments, we highlight the stability of this DyP encapsulin under harsh conditions and show that DyP catalytic activity is highest at low pH. We determine the structure of the DyP‐loaded shell and free DyP via cryo‐electron microscopy, revealing the structural basis for DyP cargo loading and peroxide preference. This work lays the foundation to further explore the substrate range and physiological functions of enterobacterial DyP encapsulins.https://doi.org/10.1002/advs.202415827DyPencapsulinenterobacteriaperoxidase
spellingShingle Natalia C. Ubilla‐Rodriguez
Michael P. Andreas
Tobias W. Giessen
Structural and Biochemical Characterization of a Widespread Enterobacterial Peroxidase Encapsulin
Advanced Science
DyP
encapsulin
enterobacteria
peroxidase
title Structural and Biochemical Characterization of a Widespread Enterobacterial Peroxidase Encapsulin
title_full Structural and Biochemical Characterization of a Widespread Enterobacterial Peroxidase Encapsulin
title_fullStr Structural and Biochemical Characterization of a Widespread Enterobacterial Peroxidase Encapsulin
title_full_unstemmed Structural and Biochemical Characterization of a Widespread Enterobacterial Peroxidase Encapsulin
title_short Structural and Biochemical Characterization of a Widespread Enterobacterial Peroxidase Encapsulin
title_sort structural and biochemical characterization of a widespread enterobacterial peroxidase encapsulin
topic DyP
encapsulin
enterobacteria
peroxidase
url https://doi.org/10.1002/advs.202415827
work_keys_str_mv AT nataliacubillarodriguez structuralandbiochemicalcharacterizationofawidespreadenterobacterialperoxidaseencapsulin
AT michaelpandreas structuralandbiochemicalcharacterizationofawidespreadenterobacterialperoxidaseencapsulin
AT tobiaswgiessen structuralandbiochemicalcharacterizationofawidespreadenterobacterialperoxidaseencapsulin