Proactive and reactive movement behaviours shape the antipredator sequence in a large herbivore

Abstract Background Prey species can display antipredator movement behaviours to reduce predation risk, including proactive responses to chronic or predictable risk, and reactive responses to acute or unpredictable risk. Thus, at any given time, prey movement choice may reflect a trade-off between p...

Full description

Saved in:
Bibliographic Details
Main Authors: Charlotte Vanderlocht, Benjamin Robira, Andrea Corradini, Simone Dal Farra, Federico Ossi, Davide Righetti, Heidi C. Hauffe, Luca Pedrotti, Francesca Cagnacci
Format: Article
Language:English
Published: BMC 2025-08-01
Series:Movement Ecology
Subjects:
Online Access:https://doi.org/10.1186/s40462-025-00584-z
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1849739635978141696
author Charlotte Vanderlocht
Benjamin Robira
Andrea Corradini
Simone Dal Farra
Federico Ossi
Davide Righetti
Heidi C. Hauffe
Luca Pedrotti
Francesca Cagnacci
author_facet Charlotte Vanderlocht
Benjamin Robira
Andrea Corradini
Simone Dal Farra
Federico Ossi
Davide Righetti
Heidi C. Hauffe
Luca Pedrotti
Francesca Cagnacci
author_sort Charlotte Vanderlocht
collection DOAJ
description Abstract Background Prey species can display antipredator movement behaviours to reduce predation risk, including proactive responses to chronic or predictable risk, and reactive responses to acute or unpredictable risk. Thus, at any given time, prey movement choice may reflect a trade-off between proaction and reaction. In previous studies, proaction and reaction have generally been considered separately, which neglects their potentially simultaneous influence on animal movement decisions and overall space use. Methods In this study, we analysed how proaction and reaction interact to shape the movements of GPS-collared red deer (Cervus elaphus) in response to hunting by humans. Using an exhaustive inventory of red deer hunting events and very high-resolution canopy cover density (LiDAR), we combined movement metric (displacement and path length) models and integrated step selection functions to investigate antipredator movement responses to lethal risk on various spatiotemporal scales, considering a dynamic landscape of risk. Results Our results show that red deer either proactively avoided areas of chronic risk, or they selected canopy cover where and when risk was predictably high. However, when risk was encountered anyway, canopy cover was no longer selected, but only modulated a reactive response along a remain-to-leave continuum. This reaction was even more evident when the environment was unfamiliar, underlining the importance of memory in such reaction patterns. Conclusions We describe how proaction and reaction fuse in an antipredator sequence of interconnected movement decisions in a large herbivore, and discuss how this result may help disentangle the ecological consequences of behavioural responses to predation. Finally, we lay the foundations for further investigations into the origins of similarities and differences between proactive and reactive movement responses.
format Article
id doaj-art-db2fb0a7afc5475f9531a8f878df84cb
institution DOAJ
issn 2051-3933
language English
publishDate 2025-08-01
publisher BMC
record_format Article
series Movement Ecology
spelling doaj-art-db2fb0a7afc5475f9531a8f878df84cb2025-08-20T03:06:13ZengBMCMovement Ecology2051-39332025-08-0113111610.1186/s40462-025-00584-zProactive and reactive movement behaviours shape the antipredator sequence in a large herbivoreCharlotte Vanderlocht0Benjamin Robira1Andrea Corradini2Simone Dal Farra3Federico Ossi4Davide Righetti5Heidi C. Hauffe6Luca Pedrotti7Francesca Cagnacci8Animal Ecology Research Unit, Research and Innovation Centre, Fondazione Edmund MachAnimal Ecology Research Unit, Research and Innovation Centre, Fondazione Edmund MachAnimal Ecology Research Unit, Research and Innovation Centre, Fondazione Edmund MachAnimal Ecology Research Unit, Research and Innovation Centre, Fondazione Edmund MachAnimal Ecology Research Unit, Research and Innovation Centre, Fondazione Edmund MachWildlife Office, Autonomous Province of Bolzano - South TyrolConservation Genomics Research Unit, Research and Innovation Centre, Fondazione Edmund MachStelvio National Park Office - Sustainable Development and Protected Areas Service, Autonomous Province of TrentoAnimal Ecology Research Unit, Research and Innovation Centre, Fondazione Edmund MachAbstract Background Prey species can display antipredator movement behaviours to reduce predation risk, including proactive responses to chronic or predictable risk, and reactive responses to acute or unpredictable risk. Thus, at any given time, prey movement choice may reflect a trade-off between proaction and reaction. In previous studies, proaction and reaction have generally been considered separately, which neglects their potentially simultaneous influence on animal movement decisions and overall space use. Methods In this study, we analysed how proaction and reaction interact to shape the movements of GPS-collared red deer (Cervus elaphus) in response to hunting by humans. Using an exhaustive inventory of red deer hunting events and very high-resolution canopy cover density (LiDAR), we combined movement metric (displacement and path length) models and integrated step selection functions to investigate antipredator movement responses to lethal risk on various spatiotemporal scales, considering a dynamic landscape of risk. Results Our results show that red deer either proactively avoided areas of chronic risk, or they selected canopy cover where and when risk was predictably high. However, when risk was encountered anyway, canopy cover was no longer selected, but only modulated a reactive response along a remain-to-leave continuum. This reaction was even more evident when the environment was unfamiliar, underlining the importance of memory in such reaction patterns. Conclusions We describe how proaction and reaction fuse in an antipredator sequence of interconnected movement decisions in a large herbivore, and discuss how this result may help disentangle the ecological consequences of behavioural responses to predation. Finally, we lay the foundations for further investigations into the origins of similarities and differences between proactive and reactive movement responses.https://doi.org/10.1186/s40462-025-00584-zPredator–prey dynamicsLandscape of fearSchedule of fearBio-loggingDisplacementPath length
spellingShingle Charlotte Vanderlocht
Benjamin Robira
Andrea Corradini
Simone Dal Farra
Federico Ossi
Davide Righetti
Heidi C. Hauffe
Luca Pedrotti
Francesca Cagnacci
Proactive and reactive movement behaviours shape the antipredator sequence in a large herbivore
Movement Ecology
Predator–prey dynamics
Landscape of fear
Schedule of fear
Bio-logging
Displacement
Path length
title Proactive and reactive movement behaviours shape the antipredator sequence in a large herbivore
title_full Proactive and reactive movement behaviours shape the antipredator sequence in a large herbivore
title_fullStr Proactive and reactive movement behaviours shape the antipredator sequence in a large herbivore
title_full_unstemmed Proactive and reactive movement behaviours shape the antipredator sequence in a large herbivore
title_short Proactive and reactive movement behaviours shape the antipredator sequence in a large herbivore
title_sort proactive and reactive movement behaviours shape the antipredator sequence in a large herbivore
topic Predator–prey dynamics
Landscape of fear
Schedule of fear
Bio-logging
Displacement
Path length
url https://doi.org/10.1186/s40462-025-00584-z
work_keys_str_mv AT charlottevanderlocht proactiveandreactivemovementbehavioursshapetheantipredatorsequenceinalargeherbivore
AT benjaminrobira proactiveandreactivemovementbehavioursshapetheantipredatorsequenceinalargeherbivore
AT andreacorradini proactiveandreactivemovementbehavioursshapetheantipredatorsequenceinalargeherbivore
AT simonedalfarra proactiveandreactivemovementbehavioursshapetheantipredatorsequenceinalargeherbivore
AT federicoossi proactiveandreactivemovementbehavioursshapetheantipredatorsequenceinalargeherbivore
AT daviderighetti proactiveandreactivemovementbehavioursshapetheantipredatorsequenceinalargeherbivore
AT heidichauffe proactiveandreactivemovementbehavioursshapetheantipredatorsequenceinalargeherbivore
AT lucapedrotti proactiveandreactivemovementbehavioursshapetheantipredatorsequenceinalargeherbivore
AT francescacagnacci proactiveandreactivemovementbehavioursshapetheantipredatorsequenceinalargeherbivore