Regulatory Divergence of Transcript Isoforms in a Mammalian Model System.

Phenotypic differences between species are driven by changes in gene expression and, by extension, by modifications in the regulation of the transcriptome. Investigation of mammalian transcriptome divergence has been restricted to analysis of bulk gene expression levels and gene-internal splicing. U...

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Main Authors: Sarah Leigh-Brown, Angela Goncalves, David Thybert, Klara Stefflova, Stephen Watt, Paul Flicek, Alvis Brazma, John C Marioni, Duncan T Odom
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2015-01-01
Series:PLoS ONE
Online Access:https://doi.org/10.1371/journal.pone.0137367
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author Sarah Leigh-Brown
Angela Goncalves
David Thybert
Klara Stefflova
Stephen Watt
Paul Flicek
Alvis Brazma
John C Marioni
Duncan T Odom
author_facet Sarah Leigh-Brown
Angela Goncalves
David Thybert
Klara Stefflova
Stephen Watt
Paul Flicek
Alvis Brazma
John C Marioni
Duncan T Odom
author_sort Sarah Leigh-Brown
collection DOAJ
description Phenotypic differences between species are driven by changes in gene expression and, by extension, by modifications in the regulation of the transcriptome. Investigation of mammalian transcriptome divergence has been restricted to analysis of bulk gene expression levels and gene-internal splicing. Using allele-specific expression analysis in inter-strain hybrids of Mus musculus, we determined the contribution of multiple cellular regulatory systems to transcriptome divergence, including: alternative promoter usage, transcription start site selection, cassette exon usage, alternative last exon usage, and alternative polyadenylation site choice. Between mouse strains, a fifth of genes have variations in isoform usage that contribute to transcriptomic changes, half of which alter encoded amino acid sequence. Virtually all divergence in isoform usage altered the post-transcriptional regulatory instructions in gene UTRs. Furthermore, most genes with isoform differences between strains contain changes originating from multiple regulatory systems. This result indicates widespread cross-talk and coordination exists among different regulatory systems. Overall, isoform usage diverges in parallel with and independently to gene expression evolution, and the cis and trans regulatory contribution to each differs significantly.
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spelling doaj-art-d8a1019b5ad84ccbbf80157fe88db89a2025-08-20T03:10:08ZengPublic Library of Science (PLoS)PLoS ONE1932-62032015-01-01109e013736710.1371/journal.pone.0137367Regulatory Divergence of Transcript Isoforms in a Mammalian Model System.Sarah Leigh-BrownAngela GoncalvesDavid ThybertKlara StefflovaStephen WattPaul FlicekAlvis BrazmaJohn C MarioniDuncan T OdomPhenotypic differences between species are driven by changes in gene expression and, by extension, by modifications in the regulation of the transcriptome. Investigation of mammalian transcriptome divergence has been restricted to analysis of bulk gene expression levels and gene-internal splicing. Using allele-specific expression analysis in inter-strain hybrids of Mus musculus, we determined the contribution of multiple cellular regulatory systems to transcriptome divergence, including: alternative promoter usage, transcription start site selection, cassette exon usage, alternative last exon usage, and alternative polyadenylation site choice. Between mouse strains, a fifth of genes have variations in isoform usage that contribute to transcriptomic changes, half of which alter encoded amino acid sequence. Virtually all divergence in isoform usage altered the post-transcriptional regulatory instructions in gene UTRs. Furthermore, most genes with isoform differences between strains contain changes originating from multiple regulatory systems. This result indicates widespread cross-talk and coordination exists among different regulatory systems. Overall, isoform usage diverges in parallel with and independently to gene expression evolution, and the cis and trans regulatory contribution to each differs significantly.https://doi.org/10.1371/journal.pone.0137367
spellingShingle Sarah Leigh-Brown
Angela Goncalves
David Thybert
Klara Stefflova
Stephen Watt
Paul Flicek
Alvis Brazma
John C Marioni
Duncan T Odom
Regulatory Divergence of Transcript Isoforms in a Mammalian Model System.
PLoS ONE
title Regulatory Divergence of Transcript Isoforms in a Mammalian Model System.
title_full Regulatory Divergence of Transcript Isoforms in a Mammalian Model System.
title_fullStr Regulatory Divergence of Transcript Isoforms in a Mammalian Model System.
title_full_unstemmed Regulatory Divergence of Transcript Isoforms in a Mammalian Model System.
title_short Regulatory Divergence of Transcript Isoforms in a Mammalian Model System.
title_sort regulatory divergence of transcript isoforms in a mammalian model system
url https://doi.org/10.1371/journal.pone.0137367
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