Trait coordination and trade-offs constrain the diversity of water use strategies in Mediterranean woody plants

Abstract The diversity of water-use strategies among dryland plants has been the focus of extensive research, but important knowledge gaps remain. Comprehensive surveys of water-use traits encompassing multiple species growing at contrasting sites are needed to further advance current understanding...

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Main Authors: Francisco J. Muñoz-Gálvez, José I. Querejeta, Cristina Moreno-Gutiérrez, Wei Ren, Enrique G. de la Riva, Iván Prieto
Format: Article
Language:English
Published: Nature Portfolio 2025-05-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-025-59348-3
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author Francisco J. Muñoz-Gálvez
José I. Querejeta
Cristina Moreno-Gutiérrez
Wei Ren
Enrique G. de la Riva
Iván Prieto
author_facet Francisco J. Muñoz-Gálvez
José I. Querejeta
Cristina Moreno-Gutiérrez
Wei Ren
Enrique G. de la Riva
Iván Prieto
author_sort Francisco J. Muñoz-Gálvez
collection DOAJ
description Abstract The diversity of water-use strategies among dryland plants has been the focus of extensive research, but important knowledge gaps remain. Comprehensive surveys of water-use traits encompassing multiple species growing at contrasting sites are needed to further advance current understanding of plant water use in drylands. Here we show that ecohydrological niche segregation driven by differences in water uptake depth among coexisting species is widespread across Mediterranean plant communities, as evidenced by soil and stem water isotopes measured in 62 native species growing at 10 sites with contrasting climatic conditions. Foliar carbon and oxygen isotopes revealed that leaf-level stomatal regulation stringency and water-use efficiency also differ markedly among coexisting species, and are both coordinated with water uptake depth. Larger and taller woody species use a greater proportion of deeper soil water, display more conservative water use traits at leaf level (“water-savers”) and show greater investment in foliage relative to shoots. Conversely, smaller species rely mainly on shallow soil water, exhibit a more profligate water use strategy (“water-spenders”) and prioritize investment in shoots over foliage. Drought stress favours coordination between above and belowground water-use traits, resulting in unavoidable trade-offs that constrain the diversity of whole-plant water use strategies in Mediterranean plant communities.
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spelling doaj-art-d76c677f28a14ae4b8395fdcdc99bf652025-08-20T02:10:50ZengNature PortfolioNature Communications2041-17232025-05-0116111710.1038/s41467-025-59348-3Trait coordination and trade-offs constrain the diversity of water use strategies in Mediterranean woody plantsFrancisco J. Muñoz-Gálvez0José I. Querejeta1Cristina Moreno-Gutiérrez2Wei Ren3Enrique G. de la Riva4Iván Prieto5Centro de Edafología y Biología Aplicada del Segura (CEBAS), Consejo Superior de Investigaciones CientíficasCentro de Edafología y Biología Aplicada del Segura (CEBAS), Consejo Superior de Investigaciones CientíficasCentro de Edafología y Biología Aplicada del Segura (CEBAS), Consejo Superior de Investigaciones CientíficasInstitute of International Rivers and Eco-Security, Yunnan UniversityÁrea de Ecología, Facultad de Ciencias Biológicas y Ambientales, Departamento de Biodiversidad y Gestión Ambiental, Universidad de LeónÁrea de Ecología, Facultad de Ciencias Biológicas y Ambientales, Departamento de Biodiversidad y Gestión Ambiental, Universidad de LeónAbstract The diversity of water-use strategies among dryland plants has been the focus of extensive research, but important knowledge gaps remain. Comprehensive surveys of water-use traits encompassing multiple species growing at contrasting sites are needed to further advance current understanding of plant water use in drylands. Here we show that ecohydrological niche segregation driven by differences in water uptake depth among coexisting species is widespread across Mediterranean plant communities, as evidenced by soil and stem water isotopes measured in 62 native species growing at 10 sites with contrasting climatic conditions. Foliar carbon and oxygen isotopes revealed that leaf-level stomatal regulation stringency and water-use efficiency also differ markedly among coexisting species, and are both coordinated with water uptake depth. Larger and taller woody species use a greater proportion of deeper soil water, display more conservative water use traits at leaf level (“water-savers”) and show greater investment in foliage relative to shoots. Conversely, smaller species rely mainly on shallow soil water, exhibit a more profligate water use strategy (“water-spenders”) and prioritize investment in shoots over foliage. Drought stress favours coordination between above and belowground water-use traits, resulting in unavoidable trade-offs that constrain the diversity of whole-plant water use strategies in Mediterranean plant communities.https://doi.org/10.1038/s41467-025-59348-3
spellingShingle Francisco J. Muñoz-Gálvez
José I. Querejeta
Cristina Moreno-Gutiérrez
Wei Ren
Enrique G. de la Riva
Iván Prieto
Trait coordination and trade-offs constrain the diversity of water use strategies in Mediterranean woody plants
Nature Communications
title Trait coordination and trade-offs constrain the diversity of water use strategies in Mediterranean woody plants
title_full Trait coordination and trade-offs constrain the diversity of water use strategies in Mediterranean woody plants
title_fullStr Trait coordination and trade-offs constrain the diversity of water use strategies in Mediterranean woody plants
title_full_unstemmed Trait coordination and trade-offs constrain the diversity of water use strategies in Mediterranean woody plants
title_short Trait coordination and trade-offs constrain the diversity of water use strategies in Mediterranean woody plants
title_sort trait coordination and trade offs constrain the diversity of water use strategies in mediterranean woody plants
url https://doi.org/10.1038/s41467-025-59348-3
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