Loss of CD98HC phosphorylation by ATM impairs antiporter trafficking and drives glutamate toxicity in Ataxia telangiectasia

Abstract Ataxia-telangiectasia is a rare genetic disorder characterized by neurological defects, immunodeficiency, cancer predisposition, radiosensitivity, decreased blood vessel integrity, and diabetes. ATM, the protein mutated in Ataxia-telangiectasia, responds to DNA damage and oxidative stress,...

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Main Authors: July Carolina Romero, Sonal S. Tonapi, Manish Parihar, Eva Loranc, Henry E. Miller, Liesl A. Lawrence, Nicklas Bassani, Daniel G. Robledo, Lin Cao, Jia Nie, Kairi Kanda, Aiola Stoja, Natalia Garcia, Aparna Gorthi, Brian J. Stoveken, Teresa W-M Fan, Teresa A. Cassel, Shan Zha, James D. Lechleiter, Nicolas Musi, Lily Q. Dong, Andrew N. Lane, Alexander J. R. Bishop
Format: Article
Language:English
Published: Nature Portfolio 2025-06-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-025-60304-4
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author July Carolina Romero
Sonal S. Tonapi
Manish Parihar
Eva Loranc
Henry E. Miller
Liesl A. Lawrence
Nicklas Bassani
Daniel G. Robledo
Lin Cao
Jia Nie
Kairi Kanda
Aiola Stoja
Natalia Garcia
Aparna Gorthi
Brian J. Stoveken
Teresa W-M Fan
Teresa A. Cassel
Shan Zha
James D. Lechleiter
Nicolas Musi
Lily Q. Dong
Andrew N. Lane
Alexander J. R. Bishop
author_facet July Carolina Romero
Sonal S. Tonapi
Manish Parihar
Eva Loranc
Henry E. Miller
Liesl A. Lawrence
Nicklas Bassani
Daniel G. Robledo
Lin Cao
Jia Nie
Kairi Kanda
Aiola Stoja
Natalia Garcia
Aparna Gorthi
Brian J. Stoveken
Teresa W-M Fan
Teresa A. Cassel
Shan Zha
James D. Lechleiter
Nicolas Musi
Lily Q. Dong
Andrew N. Lane
Alexander J. R. Bishop
author_sort July Carolina Romero
collection DOAJ
description Abstract Ataxia-telangiectasia is a rare genetic disorder characterized by neurological defects, immunodeficiency, cancer predisposition, radiosensitivity, decreased blood vessel integrity, and diabetes. ATM, the protein mutated in Ataxia-telangiectasia, responds to DNA damage and oxidative stress, but its functional relationship to the progressive clinical manifestation of this disorder is not understood. CD98HC chaperones cystine/glutamate and cationic/neutral amino acid antiporters to the cell membrane, and CD98HC phosphorylation by ATM accelerates membrane localization to acutely increase amino acid transport. Loss of ATM impacts tissues reliant on heterodimeric amino acid transporters relevant to Ataxia-telangiectasia phenotypes, such as endothelial cells (telangiectasia) and pancreatic α-cells (fatty liver and diabetes), with toxic glutamate accumulation. Bypassing the antiporters restores intracellular metabolic balance in ATM-deficient cells and mouse models. These findings provide insight into the long-known benefits of N-acetyl cysteine in Ataxia-telangiectasia cells beyond oxidative stress through removing glutamate excess by producing glutathione.
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publishDate 2025-06-01
publisher Nature Portfolio
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spelling doaj-art-d3673f3b29a8450eb8c2a45a0a22de222025-08-20T03:26:43ZengNature PortfolioNature Communications2041-17232025-06-0116112110.1038/s41467-025-60304-4Loss of CD98HC phosphorylation by ATM impairs antiporter trafficking and drives glutamate toxicity in Ataxia telangiectasiaJuly Carolina Romero0Sonal S. Tonapi1Manish Parihar2Eva Loranc3Henry E. Miller4Liesl A. Lawrence5Nicklas Bassani6Daniel G. Robledo7Lin Cao8Jia Nie9Kairi Kanda10Aiola Stoja11Natalia Garcia12Aparna Gorthi13Brian J. Stoveken14Teresa W-M Fan15Teresa A. Cassel16Shan Zha17James D. Lechleiter18Nicolas Musi19Lily Q. Dong20Andrew N. Lane21Alexander J. R. Bishop22Cell Systems and Anatomy, UT Health San AntonioCell Systems and Anatomy, UT Health San AntonioCell Systems and Anatomy, UT Health San AntonioGreehey Children’s Cancer Research Institute, UT Health San AntonioGreehey Children’s Cancer Research Institute, UT Health San AntonioCell Systems and Anatomy, UT Health San AntonioCell Systems and Anatomy, UT Health San AntonioGreehey Children’s Cancer Research Institute, UT Health San AntonioGreehey Children’s Cancer Research Institute, UT Health San AntonioBarshop Institute for Longevity and Aging Studies, Health San AntonioCell Systems and Anatomy, UT Health San AntonioCell Systems and Anatomy, UT Health San AntonioCell Systems and Anatomy, UT Health San AntonioCell Systems and Anatomy, UT Health San AntonioCell Systems and Anatomy, UT Health San AntonioToxicology and Cancer Biology, Markey Cancer Center, University of KentuckyToxicology and Cancer Biology, Markey Cancer Center, University of KentuckyInstitute for Cancer Genetics, Vagelos College of Physicians and Surgeons, Columbia UniversityCell Systems and Anatomy, UT Health San AntonioBarshop Institute for Longevity and Aging Studies, Health San AntonioCell Systems and Anatomy, UT Health San AntonioToxicology and Cancer Biology, Markey Cancer Center, University of KentuckyCell Systems and Anatomy, UT Health San AntonioAbstract Ataxia-telangiectasia is a rare genetic disorder characterized by neurological defects, immunodeficiency, cancer predisposition, radiosensitivity, decreased blood vessel integrity, and diabetes. ATM, the protein mutated in Ataxia-telangiectasia, responds to DNA damage and oxidative stress, but its functional relationship to the progressive clinical manifestation of this disorder is not understood. CD98HC chaperones cystine/glutamate and cationic/neutral amino acid antiporters to the cell membrane, and CD98HC phosphorylation by ATM accelerates membrane localization to acutely increase amino acid transport. Loss of ATM impacts tissues reliant on heterodimeric amino acid transporters relevant to Ataxia-telangiectasia phenotypes, such as endothelial cells (telangiectasia) and pancreatic α-cells (fatty liver and diabetes), with toxic glutamate accumulation. Bypassing the antiporters restores intracellular metabolic balance in ATM-deficient cells and mouse models. These findings provide insight into the long-known benefits of N-acetyl cysteine in Ataxia-telangiectasia cells beyond oxidative stress through removing glutamate excess by producing glutathione.https://doi.org/10.1038/s41467-025-60304-4
spellingShingle July Carolina Romero
Sonal S. Tonapi
Manish Parihar
Eva Loranc
Henry E. Miller
Liesl A. Lawrence
Nicklas Bassani
Daniel G. Robledo
Lin Cao
Jia Nie
Kairi Kanda
Aiola Stoja
Natalia Garcia
Aparna Gorthi
Brian J. Stoveken
Teresa W-M Fan
Teresa A. Cassel
Shan Zha
James D. Lechleiter
Nicolas Musi
Lily Q. Dong
Andrew N. Lane
Alexander J. R. Bishop
Loss of CD98HC phosphorylation by ATM impairs antiporter trafficking and drives glutamate toxicity in Ataxia telangiectasia
Nature Communications
title Loss of CD98HC phosphorylation by ATM impairs antiporter trafficking and drives glutamate toxicity in Ataxia telangiectasia
title_full Loss of CD98HC phosphorylation by ATM impairs antiporter trafficking and drives glutamate toxicity in Ataxia telangiectasia
title_fullStr Loss of CD98HC phosphorylation by ATM impairs antiporter trafficking and drives glutamate toxicity in Ataxia telangiectasia
title_full_unstemmed Loss of CD98HC phosphorylation by ATM impairs antiporter trafficking and drives glutamate toxicity in Ataxia telangiectasia
title_short Loss of CD98HC phosphorylation by ATM impairs antiporter trafficking and drives glutamate toxicity in Ataxia telangiectasia
title_sort loss of cd98hc phosphorylation by atm impairs antiporter trafficking and drives glutamate toxicity in ataxia telangiectasia
url https://doi.org/10.1038/s41467-025-60304-4
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