Host-specific sensing of coronaviruses and picornaviruses by the CARD8 inflammasome.

Hosts have evolved diverse strategies to respond to microbial infections, including the detection of pathogen-encoded proteases by inflammasome-forming sensors such as NLRP1 and CARD8. Here, we find that the 3CL protease (3CLpro) encoded by diverse coronaviruses, including Severe Acute Respiratory S...

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Main Authors: Brian V Tsu, Rimjhim Agarwal, Nandan S Gokhale, Jessie Kulsuptrakul, Andrew P Ryan, Elizabeth J Fay, Lennice K Castro, Christopher Beierschmitt, Christina Yap, Elizabeth A Turcotte, Sofia E Delgado-Rodriguez, Russell E Vance, Jennifer L Hyde, Ram Savan, Patrick S Mitchell, Matthew D Daugherty
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2023-06-01
Series:PLoS Biology
Online Access:https://doi.org/10.1371/journal.pbio.3002144
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author Brian V Tsu
Rimjhim Agarwal
Nandan S Gokhale
Jessie Kulsuptrakul
Andrew P Ryan
Elizabeth J Fay
Lennice K Castro
Christopher Beierschmitt
Christina Yap
Elizabeth A Turcotte
Sofia E Delgado-Rodriguez
Russell E Vance
Jennifer L Hyde
Ram Savan
Patrick S Mitchell
Matthew D Daugherty
author_facet Brian V Tsu
Rimjhim Agarwal
Nandan S Gokhale
Jessie Kulsuptrakul
Andrew P Ryan
Elizabeth J Fay
Lennice K Castro
Christopher Beierschmitt
Christina Yap
Elizabeth A Turcotte
Sofia E Delgado-Rodriguez
Russell E Vance
Jennifer L Hyde
Ram Savan
Patrick S Mitchell
Matthew D Daugherty
author_sort Brian V Tsu
collection DOAJ
description Hosts have evolved diverse strategies to respond to microbial infections, including the detection of pathogen-encoded proteases by inflammasome-forming sensors such as NLRP1 and CARD8. Here, we find that the 3CL protease (3CLpro) encoded by diverse coronaviruses, including Severe Acute Respiratory Syndrome Coronavirus 2 (SARS-CoV-2), cleaves a rapidly evolving region of human CARD8 and activates a robust inflammasome response. CARD8 is required for cell death and the release of pro-inflammatory cytokines during SARS-CoV-2 infection. We further find that natural variation alters CARD8 sensing of 3CLpro, including 3CLpro-mediated antagonism rather than activation of megabat CARD8. Likewise, we find that a single nucleotide polymorphism (SNP) in humans reduces CARD8's ability to sense coronavirus 3CLpros and, instead, enables sensing of 3C proteases (3Cpro) from select picornaviruses. Our findings demonstrate that CARD8 is a broad sensor of viral protease activities and suggests that CARD8 diversity contributes to inter- and intraspecies variation in inflammasome-mediated viral sensing and immunopathology.
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spelling doaj-art-cdf5043f3dfb4817b3329b51ef3b0db42025-08-20T02:38:31ZengPublic Library of Science (PLoS)PLoS Biology1544-91731545-78852023-06-01216e300214410.1371/journal.pbio.3002144Host-specific sensing of coronaviruses and picornaviruses by the CARD8 inflammasome.Brian V TsuRimjhim AgarwalNandan S GokhaleJessie KulsuptrakulAndrew P RyanElizabeth J FayLennice K CastroChristopher BeierschmittChristina YapElizabeth A TurcotteSofia E Delgado-RodriguezRussell E VanceJennifer L HydeRam SavanPatrick S MitchellMatthew D DaughertyHosts have evolved diverse strategies to respond to microbial infections, including the detection of pathogen-encoded proteases by inflammasome-forming sensors such as NLRP1 and CARD8. Here, we find that the 3CL protease (3CLpro) encoded by diverse coronaviruses, including Severe Acute Respiratory Syndrome Coronavirus 2 (SARS-CoV-2), cleaves a rapidly evolving region of human CARD8 and activates a robust inflammasome response. CARD8 is required for cell death and the release of pro-inflammatory cytokines during SARS-CoV-2 infection. We further find that natural variation alters CARD8 sensing of 3CLpro, including 3CLpro-mediated antagonism rather than activation of megabat CARD8. Likewise, we find that a single nucleotide polymorphism (SNP) in humans reduces CARD8's ability to sense coronavirus 3CLpros and, instead, enables sensing of 3C proteases (3Cpro) from select picornaviruses. Our findings demonstrate that CARD8 is a broad sensor of viral protease activities and suggests that CARD8 diversity contributes to inter- and intraspecies variation in inflammasome-mediated viral sensing and immunopathology.https://doi.org/10.1371/journal.pbio.3002144
spellingShingle Brian V Tsu
Rimjhim Agarwal
Nandan S Gokhale
Jessie Kulsuptrakul
Andrew P Ryan
Elizabeth J Fay
Lennice K Castro
Christopher Beierschmitt
Christina Yap
Elizabeth A Turcotte
Sofia E Delgado-Rodriguez
Russell E Vance
Jennifer L Hyde
Ram Savan
Patrick S Mitchell
Matthew D Daugherty
Host-specific sensing of coronaviruses and picornaviruses by the CARD8 inflammasome.
PLoS Biology
title Host-specific sensing of coronaviruses and picornaviruses by the CARD8 inflammasome.
title_full Host-specific sensing of coronaviruses and picornaviruses by the CARD8 inflammasome.
title_fullStr Host-specific sensing of coronaviruses and picornaviruses by the CARD8 inflammasome.
title_full_unstemmed Host-specific sensing of coronaviruses and picornaviruses by the CARD8 inflammasome.
title_short Host-specific sensing of coronaviruses and picornaviruses by the CARD8 inflammasome.
title_sort host specific sensing of coronaviruses and picornaviruses by the card8 inflammasome
url https://doi.org/10.1371/journal.pbio.3002144
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