Interleukin-15 response signature predicts RhCMV/SIV vaccine efficacy.

Simian immunodeficiency virus (SIV) challenge of rhesus macaques (RMs) vaccinated with strain 68-1 Rhesus Cytomegalovirus (RhCMV) vectors expressing SIV proteins (RhCMV/SIV) results in a binary outcome: stringent control and subsequent clearance of highly pathogenic SIV in ~55% of vaccinated RMs wit...

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Main Authors: Fredrik Barrenäs, Scott G Hansen, Lynn Law, Connor Driscoll, Richard R Green, Elise Smith, Jean Chang, Inah Golez, Taryn Urion, Xinxia Peng, Leanne Whitmore, Daniel Newhouse, Colette M Hughes, David Morrow, Kurt T Randall, Andrea N Selseth, Julia C Ford, Roxanne M Gilbride, Bryan E Randall, Emily Ainslie, Kelli Oswald, Rebecca Shoemaker, Randy Fast, William J Bosche, Michael K Axthelm, Yoshinori Fukazawa, George N Pavlakis, Barbara K Felber, Slim Fourati, Rafick-Pierre Sekaly, Jeffrey D Lifson, Jan Komorowski, Ewelina Kosmider, Danica Shao, Wenjun Song, Paul T Edlefsen, Louis J Picker, Michael Gale
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2021-07-01
Series:PLoS Pathogens
Online Access:https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1009278&type=printable
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author Fredrik Barrenäs
Scott G Hansen
Lynn Law
Connor Driscoll
Richard R Green
Elise Smith
Jean Chang
Inah Golez
Taryn Urion
Xinxia Peng
Leanne Whitmore
Daniel Newhouse
Colette M Hughes
David Morrow
Kurt T Randall
Andrea N Selseth
Julia C Ford
Roxanne M Gilbride
Bryan E Randall
Emily Ainslie
Kelli Oswald
Rebecca Shoemaker
Randy Fast
William J Bosche
Michael K Axthelm
Yoshinori Fukazawa
George N Pavlakis
Barbara K Felber
Slim Fourati
Rafick-Pierre Sekaly
Jeffrey D Lifson
Jan Komorowski
Ewelina Kosmider
Danica Shao
Wenjun Song
Paul T Edlefsen
Louis J Picker
Michael Gale
author_facet Fredrik Barrenäs
Scott G Hansen
Lynn Law
Connor Driscoll
Richard R Green
Elise Smith
Jean Chang
Inah Golez
Taryn Urion
Xinxia Peng
Leanne Whitmore
Daniel Newhouse
Colette M Hughes
David Morrow
Kurt T Randall
Andrea N Selseth
Julia C Ford
Roxanne M Gilbride
Bryan E Randall
Emily Ainslie
Kelli Oswald
Rebecca Shoemaker
Randy Fast
William J Bosche
Michael K Axthelm
Yoshinori Fukazawa
George N Pavlakis
Barbara K Felber
Slim Fourati
Rafick-Pierre Sekaly
Jeffrey D Lifson
Jan Komorowski
Ewelina Kosmider
Danica Shao
Wenjun Song
Paul T Edlefsen
Louis J Picker
Michael Gale
author_sort Fredrik Barrenäs
collection DOAJ
description Simian immunodeficiency virus (SIV) challenge of rhesus macaques (RMs) vaccinated with strain 68-1 Rhesus Cytomegalovirus (RhCMV) vectors expressing SIV proteins (RhCMV/SIV) results in a binary outcome: stringent control and subsequent clearance of highly pathogenic SIV in ~55% of vaccinated RMs with no protection in the remaining 45%. Although previous work indicates that unconventionally restricted, SIV-specific, effector-memory (EM)-biased CD8+ T cell responses are necessary for efficacy, the magnitude of these responses does not predict efficacy, and the basis of protection vs. non-protection in 68-1 RhCMV/SIV vector-vaccinated RMs has not been elucidated. Here, we report that 68-1 RhCMV/SIV vector administration strikingly alters the whole blood transcriptome of vaccinated RMs, with the sustained induction of specific immune-related pathways, including immune cell, toll-like receptor (TLR), inflammasome/cell death, and interleukin-15 (IL-15) signaling, significantly correlating with subsequent vaccine efficacy. Treatment of a separate RM cohort with IL-15 confirmed the central involvement of this cytokine in the protection signature, linking the major innate and adaptive immune gene expression networks that correlate with RhCMV/SIV vaccine efficacy. This change-from-baseline IL-15 response signature was also demonstrated to significantly correlate with vaccine efficacy in an independent validation cohort of vaccinated and challenged RMs. The differential IL-15 gene set response to vaccination strongly correlated with the pre-vaccination activity of this pathway, with reduced baseline expression of IL-15 response genes significantly correlating with higher vaccine-induced induction of IL-15 signaling and subsequent vaccine protection, suggesting that a robust de novo vaccine-induced IL-15 signaling response is needed to program vaccine efficacy. Thus, the RhCMV/SIV vaccine imparts a coordinated and persistent induction of innate and adaptive immune pathways featuring IL-15, a known regulator of CD8+ T cell function, that support the ability of vaccine-elicited unconventionally restricted CD8+ T cells to mediate protection against SIV challenge.
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institution Kabale University
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publishDate 2021-07-01
publisher Public Library of Science (PLoS)
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series PLoS Pathogens
spelling doaj-art-cd57c9263ff449bb8bd672c756c1208a2025-08-20T03:25:19ZengPublic Library of Science (PLoS)PLoS Pathogens1553-73661553-73742021-07-01177e100927810.1371/journal.ppat.1009278Interleukin-15 response signature predicts RhCMV/SIV vaccine efficacy.Fredrik BarrenäsScott G HansenLynn LawConnor DriscollRichard R GreenElise SmithJean ChangInah GolezTaryn UrionXinxia PengLeanne WhitmoreDaniel NewhouseColette M HughesDavid MorrowKurt T RandallAndrea N SelsethJulia C FordRoxanne M GilbrideBryan E RandallEmily AinslieKelli OswaldRebecca ShoemakerRandy FastWilliam J BoscheMichael K AxthelmYoshinori FukazawaGeorge N PavlakisBarbara K FelberSlim FouratiRafick-Pierre SekalyJeffrey D LifsonJan KomorowskiEwelina KosmiderDanica ShaoWenjun SongPaul T EdlefsenLouis J PickerMichael GaleSimian immunodeficiency virus (SIV) challenge of rhesus macaques (RMs) vaccinated with strain 68-1 Rhesus Cytomegalovirus (RhCMV) vectors expressing SIV proteins (RhCMV/SIV) results in a binary outcome: stringent control and subsequent clearance of highly pathogenic SIV in ~55% of vaccinated RMs with no protection in the remaining 45%. Although previous work indicates that unconventionally restricted, SIV-specific, effector-memory (EM)-biased CD8+ T cell responses are necessary for efficacy, the magnitude of these responses does not predict efficacy, and the basis of protection vs. non-protection in 68-1 RhCMV/SIV vector-vaccinated RMs has not been elucidated. Here, we report that 68-1 RhCMV/SIV vector administration strikingly alters the whole blood transcriptome of vaccinated RMs, with the sustained induction of specific immune-related pathways, including immune cell, toll-like receptor (TLR), inflammasome/cell death, and interleukin-15 (IL-15) signaling, significantly correlating with subsequent vaccine efficacy. Treatment of a separate RM cohort with IL-15 confirmed the central involvement of this cytokine in the protection signature, linking the major innate and adaptive immune gene expression networks that correlate with RhCMV/SIV vaccine efficacy. This change-from-baseline IL-15 response signature was also demonstrated to significantly correlate with vaccine efficacy in an independent validation cohort of vaccinated and challenged RMs. The differential IL-15 gene set response to vaccination strongly correlated with the pre-vaccination activity of this pathway, with reduced baseline expression of IL-15 response genes significantly correlating with higher vaccine-induced induction of IL-15 signaling and subsequent vaccine protection, suggesting that a robust de novo vaccine-induced IL-15 signaling response is needed to program vaccine efficacy. Thus, the RhCMV/SIV vaccine imparts a coordinated and persistent induction of innate and adaptive immune pathways featuring IL-15, a known regulator of CD8+ T cell function, that support the ability of vaccine-elicited unconventionally restricted CD8+ T cells to mediate protection against SIV challenge.https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1009278&type=printable
spellingShingle Fredrik Barrenäs
Scott G Hansen
Lynn Law
Connor Driscoll
Richard R Green
Elise Smith
Jean Chang
Inah Golez
Taryn Urion
Xinxia Peng
Leanne Whitmore
Daniel Newhouse
Colette M Hughes
David Morrow
Kurt T Randall
Andrea N Selseth
Julia C Ford
Roxanne M Gilbride
Bryan E Randall
Emily Ainslie
Kelli Oswald
Rebecca Shoemaker
Randy Fast
William J Bosche
Michael K Axthelm
Yoshinori Fukazawa
George N Pavlakis
Barbara K Felber
Slim Fourati
Rafick-Pierre Sekaly
Jeffrey D Lifson
Jan Komorowski
Ewelina Kosmider
Danica Shao
Wenjun Song
Paul T Edlefsen
Louis J Picker
Michael Gale
Interleukin-15 response signature predicts RhCMV/SIV vaccine efficacy.
PLoS Pathogens
title Interleukin-15 response signature predicts RhCMV/SIV vaccine efficacy.
title_full Interleukin-15 response signature predicts RhCMV/SIV vaccine efficacy.
title_fullStr Interleukin-15 response signature predicts RhCMV/SIV vaccine efficacy.
title_full_unstemmed Interleukin-15 response signature predicts RhCMV/SIV vaccine efficacy.
title_short Interleukin-15 response signature predicts RhCMV/SIV vaccine efficacy.
title_sort interleukin 15 response signature predicts rhcmv siv vaccine efficacy
url https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1009278&type=printable
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