Impaired ketogenesis in Leydig Cells drives testicular aging

Abstract Testicular aging commonly leads to testosterone deficiency and impaired spermatogenesis, yet the underlying mechanisms remain elusive. Here, we show that Leydig cells are particularly vulnerable to aging processes in testis. Single-cell RNA sequencing identifies the expression of Hmgcs2, th...

Full description

Saved in:
Bibliographic Details
Main Authors: Congyuan Liu, Hao Peng, Jiajie Yu, Peng Luo, Chuanfeng Xiong, Hong Chen, Hang Fan, Yuanchen Ma, Wangsheng Ou, Suyuan Zhang, Cuifeng Yang, Lerong Zhao, Yuchen Zhang, Xiaolu Guo, Qiong Ke, Tao Wang, Chunhua Deng, Weiqiang Li, Andy Peng Xiang, Kai Xia
Format: Article
Language:English
Published: Nature Portfolio 2025-05-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-025-59591-8
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1850277626795524096
author Congyuan Liu
Hao Peng
Jiajie Yu
Peng Luo
Chuanfeng Xiong
Hong Chen
Hang Fan
Yuanchen Ma
Wangsheng Ou
Suyuan Zhang
Cuifeng Yang
Lerong Zhao
Yuchen Zhang
Xiaolu Guo
Qiong Ke
Tao Wang
Chunhua Deng
Weiqiang Li
Andy Peng Xiang
Kai Xia
author_facet Congyuan Liu
Hao Peng
Jiajie Yu
Peng Luo
Chuanfeng Xiong
Hong Chen
Hang Fan
Yuanchen Ma
Wangsheng Ou
Suyuan Zhang
Cuifeng Yang
Lerong Zhao
Yuchen Zhang
Xiaolu Guo
Qiong Ke
Tao Wang
Chunhua Deng
Weiqiang Li
Andy Peng Xiang
Kai Xia
author_sort Congyuan Liu
collection DOAJ
description Abstract Testicular aging commonly leads to testosterone deficiency and impaired spermatogenesis, yet the underlying mechanisms remain elusive. Here, we show that Leydig cells are particularly vulnerable to aging processes in testis. Single-cell RNA sequencing identifies the expression of Hmgcs2, the gene encoding rate-limiting enzyme of ketogenesis, decreases significantly in Leydig cells from aged mice. Additionally, the concentrations of ketone bodies β-hydroxybutyric acid and acetoacetic acid in young testes are substantially higher than that in serum, but significantly diminish in aged testes. Silencing of Hmgcs2 in young Leydig cells drives cell senescence and accelerated testicular aging. Mechanistically, β-hydroxybutyric acid upregulates the expression of Foxo3a by facilitating histone acetylation, thereby mitigating Leydig cells senescence and promoting testosterone production. Consistently, enhanced ketogenesis by genetic manipulation or oral β-hydroxybutyric acid supplementation alleviates Leydig cells senescence and ameliorates testicular aging in aged mice. These findings highlight defective ketogenesis as a pivotal factor in testicular aging, suggesting potential therapeutic avenues for addressing age-related testicular dysfunction.
format Article
id doaj-art-ccb69b59b02f4df1a3e966d6545a55b0
institution OA Journals
issn 2041-1723
language English
publishDate 2025-05-01
publisher Nature Portfolio
record_format Article
series Nature Communications
spelling doaj-art-ccb69b59b02f4df1a3e966d6545a55b02025-08-20T01:49:48ZengNature PortfolioNature Communications2041-17232025-05-0116111910.1038/s41467-025-59591-8Impaired ketogenesis in Leydig Cells drives testicular agingCongyuan Liu0Hao Peng1Jiajie Yu2Peng Luo3Chuanfeng Xiong4Hong Chen5Hang Fan6Yuanchen Ma7Wangsheng Ou8Suyuan Zhang9Cuifeng Yang10Lerong Zhao11Yuchen Zhang12Xiaolu Guo13Qiong Ke14Tao Wang15Chunhua Deng16Weiqiang Li17Andy Peng Xiang18Kai Xia19Center for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityCenter for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityDepartment of Urology and Andrology, The First Affiliated Hospital, Sun Yat-sen UniversityReproductive Medicine Center, The First Affiliated Hospital, Sun Yat-sen University, The Key Laboratory for Reproductive Medicine of Guangdong ProvinceCenter for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityCenter for Stem Cells Translational Medicine, Shenzhen Qianhai Shekou Free Trade Zone HospitalCenter for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityCenter for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityState Key Laboratory of Ophthalmology, Zhong Shan Ophthalmic Center, Sun Yat-sen UniversityCenter for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityDepartment of Urology and Andrology, The First Affiliated Hospital, Sun Yat-sen UniversityCenter for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityCenter for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityCenter for Stem Cells Translational Medicine, Shenzhen Qianhai Shekou Free Trade Zone HospitalCenter for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityCenter for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityDepartment of Urology and Andrology, The First Affiliated Hospital, Sun Yat-sen UniversityCenter for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityCenter for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityCenter for Stem Cell Biology and Tissue Engineering, Key Laboratory for Stem Cells and Tissue Engineering, Ministry of Education, Sun Yat-sen UniversityAbstract Testicular aging commonly leads to testosterone deficiency and impaired spermatogenesis, yet the underlying mechanisms remain elusive. Here, we show that Leydig cells are particularly vulnerable to aging processes in testis. Single-cell RNA sequencing identifies the expression of Hmgcs2, the gene encoding rate-limiting enzyme of ketogenesis, decreases significantly in Leydig cells from aged mice. Additionally, the concentrations of ketone bodies β-hydroxybutyric acid and acetoacetic acid in young testes are substantially higher than that in serum, but significantly diminish in aged testes. Silencing of Hmgcs2 in young Leydig cells drives cell senescence and accelerated testicular aging. Mechanistically, β-hydroxybutyric acid upregulates the expression of Foxo3a by facilitating histone acetylation, thereby mitigating Leydig cells senescence and promoting testosterone production. Consistently, enhanced ketogenesis by genetic manipulation or oral β-hydroxybutyric acid supplementation alleviates Leydig cells senescence and ameliorates testicular aging in aged mice. These findings highlight defective ketogenesis as a pivotal factor in testicular aging, suggesting potential therapeutic avenues for addressing age-related testicular dysfunction.https://doi.org/10.1038/s41467-025-59591-8
spellingShingle Congyuan Liu
Hao Peng
Jiajie Yu
Peng Luo
Chuanfeng Xiong
Hong Chen
Hang Fan
Yuanchen Ma
Wangsheng Ou
Suyuan Zhang
Cuifeng Yang
Lerong Zhao
Yuchen Zhang
Xiaolu Guo
Qiong Ke
Tao Wang
Chunhua Deng
Weiqiang Li
Andy Peng Xiang
Kai Xia
Impaired ketogenesis in Leydig Cells drives testicular aging
Nature Communications
title Impaired ketogenesis in Leydig Cells drives testicular aging
title_full Impaired ketogenesis in Leydig Cells drives testicular aging
title_fullStr Impaired ketogenesis in Leydig Cells drives testicular aging
title_full_unstemmed Impaired ketogenesis in Leydig Cells drives testicular aging
title_short Impaired ketogenesis in Leydig Cells drives testicular aging
title_sort impaired ketogenesis in leydig cells drives testicular aging
url https://doi.org/10.1038/s41467-025-59591-8
work_keys_str_mv AT congyuanliu impairedketogenesisinleydigcellsdrivestesticularaging
AT haopeng impairedketogenesisinleydigcellsdrivestesticularaging
AT jiajieyu impairedketogenesisinleydigcellsdrivestesticularaging
AT pengluo impairedketogenesisinleydigcellsdrivestesticularaging
AT chuanfengxiong impairedketogenesisinleydigcellsdrivestesticularaging
AT hongchen impairedketogenesisinleydigcellsdrivestesticularaging
AT hangfan impairedketogenesisinleydigcellsdrivestesticularaging
AT yuanchenma impairedketogenesisinleydigcellsdrivestesticularaging
AT wangshengou impairedketogenesisinleydigcellsdrivestesticularaging
AT suyuanzhang impairedketogenesisinleydigcellsdrivestesticularaging
AT cuifengyang impairedketogenesisinleydigcellsdrivestesticularaging
AT lerongzhao impairedketogenesisinleydigcellsdrivestesticularaging
AT yuchenzhang impairedketogenesisinleydigcellsdrivestesticularaging
AT xiaoluguo impairedketogenesisinleydigcellsdrivestesticularaging
AT qiongke impairedketogenesisinleydigcellsdrivestesticularaging
AT taowang impairedketogenesisinleydigcellsdrivestesticularaging
AT chunhuadeng impairedketogenesisinleydigcellsdrivestesticularaging
AT weiqiangli impairedketogenesisinleydigcellsdrivestesticularaging
AT andypengxiang impairedketogenesisinleydigcellsdrivestesticularaging
AT kaixia impairedketogenesisinleydigcellsdrivestesticularaging