Quantitative intra-Golgi transport and organization data suggest the stable compartment nature of the Golgi

How the intra-Golgi secretory transport works remains a mystery. The cisternal progression and the stable compartment models have been proposed and are under debate. Classic cisternal progression model posits that both the intra-Golgi transport and Golgi exit of secretory cargos should occur at a co...

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Main Authors: Hieng Chiong Tie, Haiyun Wang, Divyanshu Mahajan, Hilbert Yuen In Lam, Xiuping Sun, Bing Chen, Yuguang Mu, Lei Lu
Format: Article
Language:English
Published: eLife Sciences Publications Ltd 2025-07-01
Series:eLife
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Online Access:https://elifesciences.org/articles/98582
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author Hieng Chiong Tie
Haiyun Wang
Divyanshu Mahajan
Hilbert Yuen In Lam
Xiuping Sun
Bing Chen
Yuguang Mu
Lei Lu
author_facet Hieng Chiong Tie
Haiyun Wang
Divyanshu Mahajan
Hilbert Yuen In Lam
Xiuping Sun
Bing Chen
Yuguang Mu
Lei Lu
author_sort Hieng Chiong Tie
collection DOAJ
description How the intra-Golgi secretory transport works remains a mystery. The cisternal progression and the stable compartment models have been proposed and are under debate. Classic cisternal progression model posits that both the intra-Golgi transport and Golgi exit of secretory cargos should occur at a constant velocity dictated by the cisternal progression; furthermore, COPI-mediated intra-Golgi retrograde transport is essential for maintaining the Golgi organization. Leveraging our recently developed Golgi imaging tools in nocodazole-induced Golgi ministacks, we found that the intra-Golgi transport velocity of a secretory cargo decreases during their transition from the cis to the trans-side of the Golgi, and different cargos exhibit distinct velocities even within the same cisternae. We observed a vast variation in the Golgi residence times of different cargos. Remarkably, truncation of the luminal domain causes the Golgi residence time of Tac — a standard transmembrane secretory cargo without intra-Golgi recycling signals — to extend from 16 min to a notable 3.4 hr. Additionally, when COPI-mediated intra-Golgi retrograde transport was inhibited by brefeldin A, we found that nocodazole-induced Golgi can remain stacked for over 30–60 min. Therefore, our findings challenge the classical cisternal progression model and suggest the stable compartment nature of the Golgi.
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spelling doaj-art-ccb1dd8543994fbcbfdc508c5a118e212025-08-20T03:17:44ZengeLife Sciences Publications LtdeLife2050-084X2025-07-011310.7554/eLife.98582Quantitative intra-Golgi transport and organization data suggest the stable compartment nature of the GolgiHieng Chiong Tie0https://orcid.org/0000-0003-2738-8685Haiyun Wang1https://orcid.org/0009-0003-5963-2029Divyanshu Mahajan2https://orcid.org/0000-0001-5820-6763Hilbert Yuen In Lam3https://orcid.org/0000-0001-6129-2703Xiuping Sun4Bing Chen5Yuguang Mu6Lei Lu7https://orcid.org/0000-0002-8192-1471School of Biological Sciences, Nanyang Technological University, Singapore, Singapore; Medisix Therapeutics, Singapore, SingaporeSchool of Biological Sciences, Nanyang Technological University, Singapore, Singapore; Medisix Therapeutics, Singapore, SingaporeSchool of Biological Sciences, Nanyang Technological University, Singapore, SingaporeSchool of Biological Sciences, Nanyang Technological University, Singapore, SingaporeSchool of Biological Sciences, Nanyang Technological University, Singapore, SingaporeSchool of Biological Sciences, Nanyang Technological University, Singapore, SingaporeSchool of Biological Sciences, Nanyang Technological University, Singapore, SingaporeSchool of Biological Sciences, Nanyang Technological University, Singapore, SingaporeHow the intra-Golgi secretory transport works remains a mystery. The cisternal progression and the stable compartment models have been proposed and are under debate. Classic cisternal progression model posits that both the intra-Golgi transport and Golgi exit of secretory cargos should occur at a constant velocity dictated by the cisternal progression; furthermore, COPI-mediated intra-Golgi retrograde transport is essential for maintaining the Golgi organization. Leveraging our recently developed Golgi imaging tools in nocodazole-induced Golgi ministacks, we found that the intra-Golgi transport velocity of a secretory cargo decreases during their transition from the cis to the trans-side of the Golgi, and different cargos exhibit distinct velocities even within the same cisternae. We observed a vast variation in the Golgi residence times of different cargos. Remarkably, truncation of the luminal domain causes the Golgi residence time of Tac — a standard transmembrane secretory cargo without intra-Golgi recycling signals — to extend from 16 min to a notable 3.4 hr. Additionally, when COPI-mediated intra-Golgi retrograde transport was inhibited by brefeldin A, we found that nocodazole-induced Golgi can remain stacked for over 30–60 min. Therefore, our findings challenge the classical cisternal progression model and suggest the stable compartment nature of the Golgi.https://elifesciences.org/articles/98582Golgimembrane traffickingintra-Golgi transportcisternal progression modelstable compartment model
spellingShingle Hieng Chiong Tie
Haiyun Wang
Divyanshu Mahajan
Hilbert Yuen In Lam
Xiuping Sun
Bing Chen
Yuguang Mu
Lei Lu
Quantitative intra-Golgi transport and organization data suggest the stable compartment nature of the Golgi
eLife
Golgi
membrane trafficking
intra-Golgi transport
cisternal progression model
stable compartment model
title Quantitative intra-Golgi transport and organization data suggest the stable compartment nature of the Golgi
title_full Quantitative intra-Golgi transport and organization data suggest the stable compartment nature of the Golgi
title_fullStr Quantitative intra-Golgi transport and organization data suggest the stable compartment nature of the Golgi
title_full_unstemmed Quantitative intra-Golgi transport and organization data suggest the stable compartment nature of the Golgi
title_short Quantitative intra-Golgi transport and organization data suggest the stable compartment nature of the Golgi
title_sort quantitative intra golgi transport and organization data suggest the stable compartment nature of the golgi
topic Golgi
membrane trafficking
intra-Golgi transport
cisternal progression model
stable compartment model
url https://elifesciences.org/articles/98582
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