Global m6A RNA and whole 5mC DNA methylation specifically contribute to cell replicative and premature senescence induced by extrinsic oxidative stress

ABSTRACT Hydrogen peroxide (H2O2) is a typical representative substance of environmental oxidative stress. Exogenous substances can alter epigenetic modifications from the DNA to the RNA level through oxidative stress. We investigated methylation profiles of whole RNA m6A and DNA 5mC in the epitrans...

Full description

Saved in:
Bibliographic Details
Main Authors: Chenyu Zhu, Tingting Huang, Jiaqi Fu, Min Li, Luyi Tan, Xinyu Zhang, Wenli Cheng, Caiyun Lai, Zhangying Wang, Wenji Zhang, Wenjuan Zhang
Format: Article
Language:English
Published: American Society for Microbiology 2025-07-01
Series:mSystems
Subjects:
Online Access:https://journals.asm.org/doi/10.1128/msystems.01647-24
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1849303984754393088
author Chenyu Zhu
Tingting Huang
Jiaqi Fu
Min Li
Luyi Tan
Xinyu Zhang
Wenli Cheng
Caiyun Lai
Zhangying Wang
Wenji Zhang
Wenjuan Zhang
author_facet Chenyu Zhu
Tingting Huang
Jiaqi Fu
Min Li
Luyi Tan
Xinyu Zhang
Wenli Cheng
Caiyun Lai
Zhangying Wang
Wenji Zhang
Wenjuan Zhang
author_sort Chenyu Zhu
collection DOAJ
description ABSTRACT Hydrogen peroxide (H2O2) is a typical representative substance of environmental oxidative stress. Exogenous substances can alter epigenetic modifications from the DNA to the RNA level through oxidative stress. We investigated methylation profiles of whole RNA m6A and DNA 5mC in the epitranscriptome and epigenome of human lung embryonic fibroblasts in replicative senescence and H2O2-induced premature senescence. By RNA-seq, the expression of most RNA m6A and DNA 5mC regulators was reduced in both replicative and premature senescence respectively, whereas the expression of most senescence-associated secretory phenotypes, such as SASP, was increased in premature senescence. MeRIP-seq revealed that RNA m6A methylation sites were relatively conserved in replicative and premature senescence, but that premature senescence had higher levels of m6A methylation than replicative senescence. MeDIP-seq results showed that 5mC methylation was higher in replicative senescence than in premature senescence, and the methylation peak with the largest difference appeared on chromosome 19. DO enrichment analysis indicated that RNA m6A methylation played a key role in malignant tumor regulation in replicative senescence, whereas DNA 5mC promoted malignant tumor in premature senescence. Next, to explore the interaction of RNA m6A and DNA 5mC in the senescent state, we screened common hub genes for replicative and premature senescence. Four m6A-modified target genes, namely, ASPM, CENPF, MKI67, and BLM, were all closely associated with mitosis and cell cycle regulation. In addition, we also screened 5mC target genes including CDC45, TPX2, and UBE2T.IMPORTANCERNA-seq showed that most of the m6A and 5mC regulators and the majority of SASP expression were downregulated. The m6A motif was conserved in replicative and premature senescence, and its methylation was higher in replicative senescence. The most differentially 5mC methylation peak was located on chromosome 19, and its methylation was higher in premature senescence. Gene regulation by m6A in replicative senescence and 5mC in premature senescence was enriched in malignant tumors.
format Article
id doaj-art-cb92099a38cd4c0aa484cb6f495f2c27
institution Kabale University
issn 2379-5077
language English
publishDate 2025-07-01
publisher American Society for Microbiology
record_format Article
series mSystems
spelling doaj-art-cb92099a38cd4c0aa484cb6f495f2c272025-08-20T03:55:53ZengAmerican Society for MicrobiologymSystems2379-50772025-07-0110710.1128/msystems.01647-24Global m6A RNA and whole 5mC DNA methylation specifically contribute to cell replicative and premature senescence induced by extrinsic oxidative stressChenyu Zhu0Tingting Huang1Jiaqi Fu2Min Li3Luyi Tan4Xinyu Zhang5Wenli Cheng6Caiyun Lai7Zhangying Wang8Wenji Zhang9Wenjuan Zhang10Department of Public Health and Preventive Medicine, School of Medicine, Jinan University, Guangzhou, Guangdong, ChinaDepartment of Public Health and Preventive Medicine, School of Medicine, Jinan University, Guangzhou, Guangdong, ChinaDepartment of Public Health and Preventive Medicine, School of Medicine, Jinan University, Guangzhou, Guangdong, ChinaDepartment of Public Health and Preventive Medicine, School of Medicine, Jinan University, Guangzhou, Guangdong, ChinaDepartment of Public Health and Preventive Medicine, School of Medicine, Jinan University, Guangzhou, Guangdong, ChinaDepartment of Public Health and Preventive Medicine, School of Medicine, Jinan University, Guangzhou, Guangdong, ChinaDepartment of Public Health and Preventive Medicine, School of Medicine, Jinan University, Guangzhou, Guangdong, ChinaDepartment of Public Health and Preventive Medicine, School of Medicine, Jinan University, Guangzhou, Guangdong, ChinaKey Laboratory of Crop Genetic Improvement of Guangdong Province, Crops Research Institute, Guangdong Academy of Agricultural Sciences, Guangzhou, Guangdong, ChinaKey Laboratory of Crop Genetic Improvement of Guangdong Province, Crops Research Institute, Guangdong Academy of Agricultural Sciences, Guangzhou, Guangdong, ChinaDepartment of Public Health and Preventive Medicine, School of Medicine, Jinan University, Guangzhou, Guangdong, ChinaABSTRACT Hydrogen peroxide (H2O2) is a typical representative substance of environmental oxidative stress. Exogenous substances can alter epigenetic modifications from the DNA to the RNA level through oxidative stress. We investigated methylation profiles of whole RNA m6A and DNA 5mC in the epitranscriptome and epigenome of human lung embryonic fibroblasts in replicative senescence and H2O2-induced premature senescence. By RNA-seq, the expression of most RNA m6A and DNA 5mC regulators was reduced in both replicative and premature senescence respectively, whereas the expression of most senescence-associated secretory phenotypes, such as SASP, was increased in premature senescence. MeRIP-seq revealed that RNA m6A methylation sites were relatively conserved in replicative and premature senescence, but that premature senescence had higher levels of m6A methylation than replicative senescence. MeDIP-seq results showed that 5mC methylation was higher in replicative senescence than in premature senescence, and the methylation peak with the largest difference appeared on chromosome 19. DO enrichment analysis indicated that RNA m6A methylation played a key role in malignant tumor regulation in replicative senescence, whereas DNA 5mC promoted malignant tumor in premature senescence. Next, to explore the interaction of RNA m6A and DNA 5mC in the senescent state, we screened common hub genes for replicative and premature senescence. Four m6A-modified target genes, namely, ASPM, CENPF, MKI67, and BLM, were all closely associated with mitosis and cell cycle regulation. In addition, we also screened 5mC target genes including CDC45, TPX2, and UBE2T.IMPORTANCERNA-seq showed that most of the m6A and 5mC regulators and the majority of SASP expression were downregulated. The m6A motif was conserved in replicative and premature senescence, and its methylation was higher in replicative senescence. The most differentially 5mC methylation peak was located on chromosome 19, and its methylation was higher in premature senescence. Gene regulation by m6A in replicative senescence and 5mC in premature senescence was enriched in malignant tumors.https://journals.asm.org/doi/10.1128/msystems.01647-24H2O2N6-methyladenosine5-methylcytosinepremature senescenceepigenetics
spellingShingle Chenyu Zhu
Tingting Huang
Jiaqi Fu
Min Li
Luyi Tan
Xinyu Zhang
Wenli Cheng
Caiyun Lai
Zhangying Wang
Wenji Zhang
Wenjuan Zhang
Global m6A RNA and whole 5mC DNA methylation specifically contribute to cell replicative and premature senescence induced by extrinsic oxidative stress
mSystems
H2O2
N6-methyladenosine
5-methylcytosine
premature senescence
epigenetics
title Global m6A RNA and whole 5mC DNA methylation specifically contribute to cell replicative and premature senescence induced by extrinsic oxidative stress
title_full Global m6A RNA and whole 5mC DNA methylation specifically contribute to cell replicative and premature senescence induced by extrinsic oxidative stress
title_fullStr Global m6A RNA and whole 5mC DNA methylation specifically contribute to cell replicative and premature senescence induced by extrinsic oxidative stress
title_full_unstemmed Global m6A RNA and whole 5mC DNA methylation specifically contribute to cell replicative and premature senescence induced by extrinsic oxidative stress
title_short Global m6A RNA and whole 5mC DNA methylation specifically contribute to cell replicative and premature senescence induced by extrinsic oxidative stress
title_sort global m6a rna and whole 5mc dna methylation specifically contribute to cell replicative and premature senescence induced by extrinsic oxidative stress
topic H2O2
N6-methyladenosine
5-methylcytosine
premature senescence
epigenetics
url https://journals.asm.org/doi/10.1128/msystems.01647-24
work_keys_str_mv AT chenyuzhu globalm6arnaandwhole5mcdnamethylationspecificallycontributetocellreplicativeandprematuresenescenceinducedbyextrinsicoxidativestress
AT tingtinghuang globalm6arnaandwhole5mcdnamethylationspecificallycontributetocellreplicativeandprematuresenescenceinducedbyextrinsicoxidativestress
AT jiaqifu globalm6arnaandwhole5mcdnamethylationspecificallycontributetocellreplicativeandprematuresenescenceinducedbyextrinsicoxidativestress
AT minli globalm6arnaandwhole5mcdnamethylationspecificallycontributetocellreplicativeandprematuresenescenceinducedbyextrinsicoxidativestress
AT luyitan globalm6arnaandwhole5mcdnamethylationspecificallycontributetocellreplicativeandprematuresenescenceinducedbyextrinsicoxidativestress
AT xinyuzhang globalm6arnaandwhole5mcdnamethylationspecificallycontributetocellreplicativeandprematuresenescenceinducedbyextrinsicoxidativestress
AT wenlicheng globalm6arnaandwhole5mcdnamethylationspecificallycontributetocellreplicativeandprematuresenescenceinducedbyextrinsicoxidativestress
AT caiyunlai globalm6arnaandwhole5mcdnamethylationspecificallycontributetocellreplicativeandprematuresenescenceinducedbyextrinsicoxidativestress
AT zhangyingwang globalm6arnaandwhole5mcdnamethylationspecificallycontributetocellreplicativeandprematuresenescenceinducedbyextrinsicoxidativestress
AT wenjizhang globalm6arnaandwhole5mcdnamethylationspecificallycontributetocellreplicativeandprematuresenescenceinducedbyextrinsicoxidativestress
AT wenjuanzhang globalm6arnaandwhole5mcdnamethylationspecificallycontributetocellreplicativeandprematuresenescenceinducedbyextrinsicoxidativestress