Backsplicing of the HIV-1 transcript generates multiple circRNAs to promote viral replication

Abstract Circular RNAs (circRNAs) are a family of non-coding RNAs that originate from a non-canonical splicing event (backsplicing) that forms covalently closed continuous loops. An analysis of the human immunodeficiency type 1 virus (HIV-1) complex splicing pattern indicated that the virus had the...

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Main Authors: Christopher Mauer, Sean Paz, Massimo Caputi
Format: Article
Language:English
Published: Nature Portfolio 2025-03-01
Series:npj Viruses
Online Access:https://doi.org/10.1038/s44298-025-00105-0
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author Christopher Mauer
Sean Paz
Massimo Caputi
author_facet Christopher Mauer
Sean Paz
Massimo Caputi
author_sort Christopher Mauer
collection DOAJ
description Abstract Circular RNAs (circRNAs) are a family of non-coding RNAs that originate from a non-canonical splicing event (backsplicing) that forms covalently closed continuous loops. An analysis of the human immunodeficiency type 1 virus (HIV-1) complex splicing pattern indicated that the virus had the potential to generate at least 15 distinct circRNAs. The predicted HIV circRNAs were amplified utilizing divergent PCR primers and confirmed by RNase R digestion and sequencing. A predictive circRNA-miRNA interaction modeling approach and a series of validation assays determined that two cellular miRNAs, miR-6727-3p and miR-4722-3p, functionally interact with a sequence present in 8 of the HIV circRNAs. Expression of miR-6727-3p and miR-4722-3p restricted HIV-1 replication while a circRNA containing the sequence recognized by miR-6727-3p and miR-4722-3p increased the production of infective virions. Additionally, miR-6727-3p and miR-4722-3p expression was upregulated following HIV-1 infection of primary CD4+ T cells. Overall, the data presented shows that HIV-1 generates circRNAs which promote viral replication by sequestering and inhibiting the functions of miR-6727-3p and miR-4722-3p.
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spelling doaj-art-c90bdeb51bcf49b395bd0fe2d3630e412025-08-20T02:49:31ZengNature Portfolionpj Viruses2948-17672025-03-013111110.1038/s44298-025-00105-0Backsplicing of the HIV-1 transcript generates multiple circRNAs to promote viral replicationChristopher Mauer0Sean Paz1Massimo Caputi2Department of Biomedical Science, Charles E. Schmidt College of Medicine, Florida Atlantic UniversityDepartment of Biomedical Science, Charles E. Schmidt College of Medicine, Florida Atlantic UniversityDepartment of Biomedical Science, Charles E. Schmidt College of Medicine, Florida Atlantic UniversityAbstract Circular RNAs (circRNAs) are a family of non-coding RNAs that originate from a non-canonical splicing event (backsplicing) that forms covalently closed continuous loops. An analysis of the human immunodeficiency type 1 virus (HIV-1) complex splicing pattern indicated that the virus had the potential to generate at least 15 distinct circRNAs. The predicted HIV circRNAs were amplified utilizing divergent PCR primers and confirmed by RNase R digestion and sequencing. A predictive circRNA-miRNA interaction modeling approach and a series of validation assays determined that two cellular miRNAs, miR-6727-3p and miR-4722-3p, functionally interact with a sequence present in 8 of the HIV circRNAs. Expression of miR-6727-3p and miR-4722-3p restricted HIV-1 replication while a circRNA containing the sequence recognized by miR-6727-3p and miR-4722-3p increased the production of infective virions. Additionally, miR-6727-3p and miR-4722-3p expression was upregulated following HIV-1 infection of primary CD4+ T cells. Overall, the data presented shows that HIV-1 generates circRNAs which promote viral replication by sequestering and inhibiting the functions of miR-6727-3p and miR-4722-3p.https://doi.org/10.1038/s44298-025-00105-0
spellingShingle Christopher Mauer
Sean Paz
Massimo Caputi
Backsplicing of the HIV-1 transcript generates multiple circRNAs to promote viral replication
npj Viruses
title Backsplicing of the HIV-1 transcript generates multiple circRNAs to promote viral replication
title_full Backsplicing of the HIV-1 transcript generates multiple circRNAs to promote viral replication
title_fullStr Backsplicing of the HIV-1 transcript generates multiple circRNAs to promote viral replication
title_full_unstemmed Backsplicing of the HIV-1 transcript generates multiple circRNAs to promote viral replication
title_short Backsplicing of the HIV-1 transcript generates multiple circRNAs to promote viral replication
title_sort backsplicing of the hiv 1 transcript generates multiple circrnas to promote viral replication
url https://doi.org/10.1038/s44298-025-00105-0
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AT massimocaputi backsplicingofthehiv1transcriptgeneratesmultiplecircrnastopromoteviralreplication