Discovery of a lipid synthesising organ in the auditory system of an insect.

Weta possess typical Ensifera ears. Each ear comprises three functional parts: two equally sized tympanal membranes, an underlying system of modified tracheal chambers, and the auditory sensory organ, the crista acustica. This organ sits within an enclosed fluid-filled channel-previously presumed to...

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Main Authors: Kathryn F Lomas, David R Greenwood, James F C Windmill, Joseph C Jackson, Jeremy Corfield, Stuart Parsons
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2012-01-01
Series:PLoS ONE
Online Access:https://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0051486&type=printable
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author Kathryn F Lomas
David R Greenwood
James F C Windmill
Joseph C Jackson
Jeremy Corfield
Stuart Parsons
author_facet Kathryn F Lomas
David R Greenwood
James F C Windmill
Joseph C Jackson
Jeremy Corfield
Stuart Parsons
author_sort Kathryn F Lomas
collection DOAJ
description Weta possess typical Ensifera ears. Each ear comprises three functional parts: two equally sized tympanal membranes, an underlying system of modified tracheal chambers, and the auditory sensory organ, the crista acustica. This organ sits within an enclosed fluid-filled channel-previously presumed to be hemolymph. The role this channel plays in insect hearing is unknown. We discovered that the fluid within the channel is not actually hemolymph, but a medium composed principally of lipid from a new class. Three-dimensional imaging of this lipid channel revealed a previously undescribed tissue structure within the channel, which we refer to as the olivarius organ. Investigations into the function of the olivarius reveal de novo lipid synthesis indicating that it is producing these lipids in situ from acetate. The auditory role of this lipid channel was investigated using Laser Doppler vibrometry of the tympanal membrane, which shows that the displacement of the membrane is significantly increased when the lipid is removed from the auditory system. Neural sensitivity of the system, however, decreased upon removal of the lipid-a surprising result considering that in a typical auditory system both the mechanical and auditory sensitivity are positively correlated. These two results coupled with 3D modelling of the auditory system lead us to hypothesize a model for weta audition, relying strongly on the presence of the lipid channel. This is the first instance of lipids being associated with an auditory system outside of the Odentocete cetaceans, demonstrating convergence for the use of lipids in hearing.
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spelling doaj-art-c7ef77ac2ed947d88b305f8b61d988f52025-08-20T02:30:56ZengPublic Library of Science (PLoS)PLoS ONE1932-62032012-01-01712e5148610.1371/journal.pone.0051486Discovery of a lipid synthesising organ in the auditory system of an insect.Kathryn F LomasDavid R GreenwoodJames F C WindmillJoseph C JacksonJeremy CorfieldStuart ParsonsWeta possess typical Ensifera ears. Each ear comprises three functional parts: two equally sized tympanal membranes, an underlying system of modified tracheal chambers, and the auditory sensory organ, the crista acustica. This organ sits within an enclosed fluid-filled channel-previously presumed to be hemolymph. The role this channel plays in insect hearing is unknown. We discovered that the fluid within the channel is not actually hemolymph, but a medium composed principally of lipid from a new class. Three-dimensional imaging of this lipid channel revealed a previously undescribed tissue structure within the channel, which we refer to as the olivarius organ. Investigations into the function of the olivarius reveal de novo lipid synthesis indicating that it is producing these lipids in situ from acetate. The auditory role of this lipid channel was investigated using Laser Doppler vibrometry of the tympanal membrane, which shows that the displacement of the membrane is significantly increased when the lipid is removed from the auditory system. Neural sensitivity of the system, however, decreased upon removal of the lipid-a surprising result considering that in a typical auditory system both the mechanical and auditory sensitivity are positively correlated. These two results coupled with 3D modelling of the auditory system lead us to hypothesize a model for weta audition, relying strongly on the presence of the lipid channel. This is the first instance of lipids being associated with an auditory system outside of the Odentocete cetaceans, demonstrating convergence for the use of lipids in hearing.https://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0051486&type=printable
spellingShingle Kathryn F Lomas
David R Greenwood
James F C Windmill
Joseph C Jackson
Jeremy Corfield
Stuart Parsons
Discovery of a lipid synthesising organ in the auditory system of an insect.
PLoS ONE
title Discovery of a lipid synthesising organ in the auditory system of an insect.
title_full Discovery of a lipid synthesising organ in the auditory system of an insect.
title_fullStr Discovery of a lipid synthesising organ in the auditory system of an insect.
title_full_unstemmed Discovery of a lipid synthesising organ in the auditory system of an insect.
title_short Discovery of a lipid synthesising organ in the auditory system of an insect.
title_sort discovery of a lipid synthesising organ in the auditory system of an insect
url https://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0051486&type=printable
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