Reawakening inflammation in the chronically injured spinal cord using lipopolysaccharide induces diverse microglial states

Abstract Background Rehabilitative training is an effective method to promote recovery following spinal cord injury (SCI), with lower training efficacy observed in the chronic stage. The increased training efficacy during the subacute period is associated with a shift towards a more adaptive or pror...

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Main Authors: Rebecca K. John, Sadie P. Vogel, Sameera Zia, Kelly V. Lee, Antoinette T. Nguyen, Abel Torres-Espin, Keith K. Fenrich, Carmen Ng, Emma K. A. Schmidt, Romana Vavrek, Pamela J. F. Raposo, Keira Smith, Karim Fouad, Jason R. Plemel
Format: Article
Language:English
Published: BMC 2025-02-01
Series:Journal of Neuroinflammation
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Online Access:https://doi.org/10.1186/s12974-025-03379-6
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author Rebecca K. John
Sadie P. Vogel
Sameera Zia
Kelly V. Lee
Antoinette T. Nguyen
Abel Torres-Espin
Keith K. Fenrich
Carmen Ng
Emma K. A. Schmidt
Romana Vavrek
Pamela J. F. Raposo
Keira Smith
Karim Fouad
Jason R. Plemel
author_facet Rebecca K. John
Sadie P. Vogel
Sameera Zia
Kelly V. Lee
Antoinette T. Nguyen
Abel Torres-Espin
Keith K. Fenrich
Carmen Ng
Emma K. A. Schmidt
Romana Vavrek
Pamela J. F. Raposo
Keira Smith
Karim Fouad
Jason R. Plemel
author_sort Rebecca K. John
collection DOAJ
description Abstract Background Rehabilitative training is an effective method to promote recovery following spinal cord injury (SCI), with lower training efficacy observed in the chronic stage. The increased training efficacy during the subacute period is associated with a shift towards a more adaptive or proreparative state induced by the SCI. A potential link is SCI-induced inflammation, which is elevated in the subacute period, and, as injection of lipopolysaccharide (LPS) alongside training improves recovery in chronic SCI, suggesting LPS could reopen a window of plasticity late after injury. Microglia may play a role in LPS-mediated plasticity as they react to LPS and are implicated in facilitating recovery following SCI. However, it is unknown how microglia change in response to LPS following SCI to promote neuroplasticity. Main body Here we used single-cell RNA sequencing to examine microglial responses in subacute and chronic SCI with and without an LPS injection. We show that subacute SCI is characterized by a disease-associated microglial (DAM) signature, while chronic SCI is highly heterogeneous, with both injury-induced and homeostatic states. DAM states exhibit predicted metabolic pathway activity and neuronal interactions that are associated with potential mediators of plasticity. With LPS injection, microglia shifted away from the homeostatic signature to a primed, translation-associated state and increased DAM in degenerated tracts caudal to the injury. Conclusion Microglial states following an inflammatory stimulus in chronic injury incompletely recapitulate the subacute injury environment, showing both overlapping and distinct microglial signatures across time and with LPS injection. Our results contribute to an understanding of how microglia and LPS-induced neuroinflammation contribute to plasticity following SCI.
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spelling doaj-art-c3d9c78cdbeb48718bb2b963d39c86882025-08-20T02:16:45ZengBMCJournal of Neuroinflammation1742-20942025-02-0122111710.1186/s12974-025-03379-6Reawakening inflammation in the chronically injured spinal cord using lipopolysaccharide induces diverse microglial statesRebecca K. John0Sadie P. Vogel1Sameera Zia2Kelly V. Lee3Antoinette T. Nguyen4Abel Torres-Espin5Keith K. Fenrich6Carmen Ng7Emma K. A. Schmidt8Romana Vavrek9Pamela J. F. Raposo10Keira Smith11Karim Fouad12Jason R. Plemel13Neuroscience and Mental Health Institute, University of AlbertaNeuroscience and Mental Health Institute, University of AlbertaNeuroscience and Mental Health Institute, University of AlbertaDepartment of Medicine, Division of Neurology, University of AlbertaDepartment of Physical Therapy, Faculty of Rehabilitation Medicine, University of AlbertaDepartment of Physical Therapy, Faculty of Rehabilitation Medicine, University of AlbertaNeuroscience and Mental Health Institute, University of AlbertaNeuroscience and Mental Health Institute, University of AlbertaNeuroscience and Mental Health Institute, University of AlbertaDepartment of Physical Therapy, Faculty of Rehabilitation Medicine, University of AlbertaDepartment of Physical Therapy, Faculty of Rehabilitation Medicine, University of AlbertaNeuroscience and Mental Health Institute, University of AlbertaNeuroscience and Mental Health Institute, University of AlbertaNeuroscience and Mental Health Institute, University of AlbertaAbstract Background Rehabilitative training is an effective method to promote recovery following spinal cord injury (SCI), with lower training efficacy observed in the chronic stage. The increased training efficacy during the subacute period is associated with a shift towards a more adaptive or proreparative state induced by the SCI. A potential link is SCI-induced inflammation, which is elevated in the subacute period, and, as injection of lipopolysaccharide (LPS) alongside training improves recovery in chronic SCI, suggesting LPS could reopen a window of plasticity late after injury. Microglia may play a role in LPS-mediated plasticity as they react to LPS and are implicated in facilitating recovery following SCI. However, it is unknown how microglia change in response to LPS following SCI to promote neuroplasticity. Main body Here we used single-cell RNA sequencing to examine microglial responses in subacute and chronic SCI with and without an LPS injection. We show that subacute SCI is characterized by a disease-associated microglial (DAM) signature, while chronic SCI is highly heterogeneous, with both injury-induced and homeostatic states. DAM states exhibit predicted metabolic pathway activity and neuronal interactions that are associated with potential mediators of plasticity. With LPS injection, microglia shifted away from the homeostatic signature to a primed, translation-associated state and increased DAM in degenerated tracts caudal to the injury. Conclusion Microglial states following an inflammatory stimulus in chronic injury incompletely recapitulate the subacute injury environment, showing both overlapping and distinct microglial signatures across time and with LPS injection. Our results contribute to an understanding of how microglia and LPS-induced neuroinflammation contribute to plasticity following SCI.https://doi.org/10.1186/s12974-025-03379-6MicrogliaSpinal cord injuryInflammationSingle-cell RNA sequencingPlasticityLipopolysaccharide
spellingShingle Rebecca K. John
Sadie P. Vogel
Sameera Zia
Kelly V. Lee
Antoinette T. Nguyen
Abel Torres-Espin
Keith K. Fenrich
Carmen Ng
Emma K. A. Schmidt
Romana Vavrek
Pamela J. F. Raposo
Keira Smith
Karim Fouad
Jason R. Plemel
Reawakening inflammation in the chronically injured spinal cord using lipopolysaccharide induces diverse microglial states
Journal of Neuroinflammation
Microglia
Spinal cord injury
Inflammation
Single-cell RNA sequencing
Plasticity
Lipopolysaccharide
title Reawakening inflammation in the chronically injured spinal cord using lipopolysaccharide induces diverse microglial states
title_full Reawakening inflammation in the chronically injured spinal cord using lipopolysaccharide induces diverse microglial states
title_fullStr Reawakening inflammation in the chronically injured spinal cord using lipopolysaccharide induces diverse microglial states
title_full_unstemmed Reawakening inflammation in the chronically injured spinal cord using lipopolysaccharide induces diverse microglial states
title_short Reawakening inflammation in the chronically injured spinal cord using lipopolysaccharide induces diverse microglial states
title_sort reawakening inflammation in the chronically injured spinal cord using lipopolysaccharide induces diverse microglial states
topic Microglia
Spinal cord injury
Inflammation
Single-cell RNA sequencing
Plasticity
Lipopolysaccharide
url https://doi.org/10.1186/s12974-025-03379-6
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