Infection and Genomic Properties of Single- and Double-Stranded DNA <i>Cellulophaga</i> Phages

Bacterial viruses (phages) are abundant and ecologically impactful, but laboratory-based experimental model systems vastly under-represent known phage diversity, particularly for ssDNA phages. Here, we characterize the genomes and infection properties of two unrelated marine flavophages—ssDNA genera...

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Main Authors: Cristina Howard-Varona, Natalie E. Solonenko, Marie Burris, Marion Urvoy, Courtney M. Sanderson, Bejamin Bolduc, Matthew B. Sullivan
Format: Article
Language:English
Published: MDPI AG 2025-03-01
Series:Viruses
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Online Access:https://www.mdpi.com/1999-4915/17/3/365
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author Cristina Howard-Varona
Natalie E. Solonenko
Marie Burris
Marion Urvoy
Courtney M. Sanderson
Bejamin Bolduc
Matthew B. Sullivan
author_facet Cristina Howard-Varona
Natalie E. Solonenko
Marie Burris
Marion Urvoy
Courtney M. Sanderson
Bejamin Bolduc
Matthew B. Sullivan
author_sort Cristina Howard-Varona
collection DOAJ
description Bacterial viruses (phages) are abundant and ecologically impactful, but laboratory-based experimental model systems vastly under-represent known phage diversity, particularly for ssDNA phages. Here, we characterize the genomes and infection properties of two unrelated marine flavophages—ssDNA generalist phage phi18:4 (6.5 Kbp) and dsDNA specialist phage phi18:1 (39.2 Kbp)—when infecting the same <i>Cellulophaga baltica</i> strain #18 (Cba18), of the class <i>Flavobacteriia</i>. Phage phi18:4 belongs to a new family of ssDNA phages, has an internal lipid membrane, and its genome encodes primarily structural proteins, as well as a DNA replication protein common to ssDNA phages and a unique lysis protein. Phage phi18:1 is a siphovirus that encodes several virulence genes, despite not having a known temperate lifestyle, a CAZy enzyme likely for regulatory purposes, and four DNA methyltransferases dispersed throughout the genome that suggest both host modulation and phage DNA protection against host restriction. Physiologically, ssDNA phage phi18:4 has a shorter latent period and smaller burst size than dsDNA phage phi18:1, and both phages efficiently infect this host. These results help augment the diversity of characterized environmental phage–host model systems by studying infections of genomically diverse phages (ssDNA vs. dsDNA) on the same host.
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spelling doaj-art-c0ba4c3f41804b5f857700acd1e3349e2025-08-20T01:50:07ZengMDPI AGViruses1999-49152025-03-0117336510.3390/v17030365Infection and Genomic Properties of Single- and Double-Stranded DNA <i>Cellulophaga</i> PhagesCristina Howard-Varona0Natalie E. Solonenko1Marie Burris2Marion Urvoy3Courtney M. Sanderson4Bejamin Bolduc5Matthew B. Sullivan6Department of Microbiology, The Ohio State University, 484 W 12th Ave, Columbus, OH 43210, USADepartment of Microbiology, The Ohio State University, 484 W 12th Ave, Columbus, OH 43210, USADepartment of Microbiology, The Ohio State University, 484 W 12th Ave, Columbus, OH 43210, USADepartment of Microbiology, The Ohio State University, 484 W 12th Ave, Columbus, OH 43210, USADepartment of Microbiology, The Ohio State University, 484 W 12th Ave, Columbus, OH 43210, USADepartment of Microbiology, The Ohio State University, 484 W 12th Ave, Columbus, OH 43210, USADepartment of Microbiology, The Ohio State University, 484 W 12th Ave, Columbus, OH 43210, USABacterial viruses (phages) are abundant and ecologically impactful, but laboratory-based experimental model systems vastly under-represent known phage diversity, particularly for ssDNA phages. Here, we characterize the genomes and infection properties of two unrelated marine flavophages—ssDNA generalist phage phi18:4 (6.5 Kbp) and dsDNA specialist phage phi18:1 (39.2 Kbp)—when infecting the same <i>Cellulophaga baltica</i> strain #18 (Cba18), of the class <i>Flavobacteriia</i>. Phage phi18:4 belongs to a new family of ssDNA phages, has an internal lipid membrane, and its genome encodes primarily structural proteins, as well as a DNA replication protein common to ssDNA phages and a unique lysis protein. Phage phi18:1 is a siphovirus that encodes several virulence genes, despite not having a known temperate lifestyle, a CAZy enzyme likely for regulatory purposes, and four DNA methyltransferases dispersed throughout the genome that suggest both host modulation and phage DNA protection against host restriction. Physiologically, ssDNA phage phi18:4 has a shorter latent period and smaller burst size than dsDNA phage phi18:1, and both phages efficiently infect this host. These results help augment the diversity of characterized environmental phage–host model systems by studying infections of genomically diverse phages (ssDNA vs. dsDNA) on the same host.https://www.mdpi.com/1999-4915/17/3/365virusbacteriophageflavophagessDNA phagedsDNA phagebacteria
spellingShingle Cristina Howard-Varona
Natalie E. Solonenko
Marie Burris
Marion Urvoy
Courtney M. Sanderson
Bejamin Bolduc
Matthew B. Sullivan
Infection and Genomic Properties of Single- and Double-Stranded DNA <i>Cellulophaga</i> Phages
Viruses
virus
bacteriophage
flavophage
ssDNA phage
dsDNA phage
bacteria
title Infection and Genomic Properties of Single- and Double-Stranded DNA <i>Cellulophaga</i> Phages
title_full Infection and Genomic Properties of Single- and Double-Stranded DNA <i>Cellulophaga</i> Phages
title_fullStr Infection and Genomic Properties of Single- and Double-Stranded DNA <i>Cellulophaga</i> Phages
title_full_unstemmed Infection and Genomic Properties of Single- and Double-Stranded DNA <i>Cellulophaga</i> Phages
title_short Infection and Genomic Properties of Single- and Double-Stranded DNA <i>Cellulophaga</i> Phages
title_sort infection and genomic properties of single and double stranded dna i cellulophaga i phages
topic virus
bacteriophage
flavophage
ssDNA phage
dsDNA phage
bacteria
url https://www.mdpi.com/1999-4915/17/3/365
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