Investigation of the acetic acid stress response in Saccharomyces cerevisiae with mutated H3 residues

Enhanced levels of acetic acid reduce the activity of yeast strains employed for industrial fermentation-based applications. Therefore, unraveling the genetic factors underlying the regulation of the tolerance and sensitivity of yeast towards acetic acid is imperative for optimising various industri...

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Main Authors: Nitu Saha, Swati Swagatika, Raghuvir Singh Tomar
Format: Article
Language:English
Published: Shared Science Publishers OG 2023-08-01
Series:Microbial Cell
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Online Access:http://microbialcell.com/researcharticles/2023a-saha-microbial-cell/
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author Nitu Saha
Swati Swagatika
Raghuvir Singh Tomar
author_facet Nitu Saha
Swati Swagatika
Raghuvir Singh Tomar
author_sort Nitu Saha
collection DOAJ
description Enhanced levels of acetic acid reduce the activity of yeast strains employed for industrial fermentation-based applications. Therefore, unraveling the genetic factors underlying the regulation of the tolerance and sensitivity of yeast towards acetic acid is imperative for optimising various industrial processes. In this communication, we have attempted to decipher the acetic acid stress response of the previously reported acetic acid-sensitive histone mutants. Revalidation using spot-test assays and growth curves revealed that five of these mutants, viz., H3K18Q, H3S28A, H3K42Q, H3Q68A, and H3F104A, are most sensitive towards the tested acetic acid concentrations. These mutants demonstrated enhanced acetic acid stress response as evidenced by the increased expression levels of AIF1, reactive oxygen species (ROS) generation, chromatin fragmentation, and aggregated actin cytoskeleton. Additionally, the mutants exhibited active cell wall damage response upon acetic acid treatment, as demonstrated by increased Slt2-phosphorylation and expression of cell wall integrity genes. Interestingly, the mutants demonstrated increased sensitivity to cell wall stress-causing agents. Finally, screening of histone H3 N-terminal tail truncation mutants revealed that the tail truncations exhibit general sensitivity to acetic acid stress. Some of these N-terminal tail truncation mutants viz., H3 [del 1-24], H3 [del 1-28], H3 [del 9-24], and H3 [del 25-36] are also sensitive to cell wall stress agents such as Congo red and caffeine suggesting that their enhanced acetic acid sensitivity may be due to cell wall stress induced by acetic acid.
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spelling doaj-art-bf73eb38a3e945b29d30f75a4a31a2b02025-08-20T02:04:34ZengShared Science Publishers OGMicrobial Cell2311-26382023-08-01101021723210.15698/mic2023.10.806Investigation of the acetic acid stress response in Saccharomyces cerevisiae with mutated H3 residuesNitu Saha0Swati Swagatika1Raghuvir Singh Tomar2Laboratory of Chromatin Biology, Department of Biological Sciences, Indian Institute of Science Education and Research Bhopal, 462066, Madhya Pradesh, India.Laboratory of Chromatin Biology, Department of Biological Sciences, Indian Institute of Science Education and Research Bhopal, 462066, Madhya Pradesh, India.Laboratory of Chromatin Biology, Department of Biological Sciences, Indian Institute of Science Education and Research Bhopal, 462066, Madhya Pradesh, India.Enhanced levels of acetic acid reduce the activity of yeast strains employed for industrial fermentation-based applications. Therefore, unraveling the genetic factors underlying the regulation of the tolerance and sensitivity of yeast towards acetic acid is imperative for optimising various industrial processes. In this communication, we have attempted to decipher the acetic acid stress response of the previously reported acetic acid-sensitive histone mutants. Revalidation using spot-test assays and growth curves revealed that five of these mutants, viz., H3K18Q, H3S28A, H3K42Q, H3Q68A, and H3F104A, are most sensitive towards the tested acetic acid concentrations. These mutants demonstrated enhanced acetic acid stress response as evidenced by the increased expression levels of AIF1, reactive oxygen species (ROS) generation, chromatin fragmentation, and aggregated actin cytoskeleton. Additionally, the mutants exhibited active cell wall damage response upon acetic acid treatment, as demonstrated by increased Slt2-phosphorylation and expression of cell wall integrity genes. Interestingly, the mutants demonstrated increased sensitivity to cell wall stress-causing agents. Finally, screening of histone H3 N-terminal tail truncation mutants revealed that the tail truncations exhibit general sensitivity to acetic acid stress. Some of these N-terminal tail truncation mutants viz., H3 [del 1-24], H3 [del 1-28], H3 [del 9-24], and H3 [del 25-36] are also sensitive to cell wall stress agents such as Congo red and caffeine suggesting that their enhanced acetic acid sensitivity may be due to cell wall stress induced by acetic acid.http://microbialcell.com/researcharticles/2023a-saha-microbial-cell/acetic acidh3 point mutantsh3 n-terminal tail truncation mutantsaif1reactive oxygen species
spellingShingle Nitu Saha
Swati Swagatika
Raghuvir Singh Tomar
Investigation of the acetic acid stress response in Saccharomyces cerevisiae with mutated H3 residues
Microbial Cell
acetic acid
h3 point mutants
h3 n-terminal tail truncation mutants
aif1
reactive oxygen species
title Investigation of the acetic acid stress response in Saccharomyces cerevisiae with mutated H3 residues
title_full Investigation of the acetic acid stress response in Saccharomyces cerevisiae with mutated H3 residues
title_fullStr Investigation of the acetic acid stress response in Saccharomyces cerevisiae with mutated H3 residues
title_full_unstemmed Investigation of the acetic acid stress response in Saccharomyces cerevisiae with mutated H3 residues
title_short Investigation of the acetic acid stress response in Saccharomyces cerevisiae with mutated H3 residues
title_sort investigation of the acetic acid stress response in saccharomyces cerevisiae with mutated h3 residues
topic acetic acid
h3 point mutants
h3 n-terminal tail truncation mutants
aif1
reactive oxygen species
url http://microbialcell.com/researcharticles/2023a-saha-microbial-cell/
work_keys_str_mv AT nitusaha investigationoftheaceticacidstressresponseinsaccharomycescerevisiaewithmutatedh3residues
AT swatiswagatika investigationoftheaceticacidstressresponseinsaccharomycescerevisiaewithmutatedh3residues
AT raghuvirsinghtomar investigationoftheaceticacidstressresponseinsaccharomycescerevisiaewithmutatedh3residues