Coiled-coil-domain-mediated TRIM E3 protein condensation modulates enzymatic activity and functional specificity

Summary: Tripartite motif (TRIM) proteins are E3 ligases with modular structures involved in diverse cellular functions. Whether biomolecular condensation broadly regulates TRIM proteins remains unclear. Here, we systematically examine 75 TRIMs in mammalian cells and find that the majority can form...

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Main Authors: Qi Chen, Peiguo Yang
Format: Article
Language:English
Published: Elsevier 2025-06-01
Series:Cell Reports
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Online Access:http://www.sciencedirect.com/science/article/pii/S2211124725005376
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author Qi Chen
Peiguo Yang
author_facet Qi Chen
Peiguo Yang
author_sort Qi Chen
collection DOAJ
description Summary: Tripartite motif (TRIM) proteins are E3 ligases with modular structures involved in diverse cellular functions. Whether biomolecular condensation broadly regulates TRIM proteins remains unclear. Here, we systematically examine 75 TRIMs in mammalian cells and find that the majority can form condensates, either individually or together with other TRIMs. The coiled-coil (CC) domain is essential for TRIM condensation, and disease-related single-nucleotide polymorphisms (SNPs) in this domain impair condensate formation. We show that condensation modulates E3 ligase activity of several TRIMs in a context-dependent manner using cellular auto-ubiquitination and biotinylated-ubiquitination assays. Proteomic analysis reveals that proteins within 21 TRIM condensates are linked to key cellular pathways and diseases. Functional screening uncovers that certain TRIM condensates regulate centriolar satellite organization, cilia assembly, and microtubule stabilization. Our work provides foundational insights into the condensation and functions of TRIM family proteins and establishes a framework for studying co-condensation specificity among highly homologous proteins.
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spelling doaj-art-bf7119fb74b34963bc14952a8dcf37962025-08-20T02:00:51ZengElsevierCell Reports2211-12472025-06-0144611576610.1016/j.celrep.2025.115766Coiled-coil-domain-mediated TRIM E3 protein condensation modulates enzymatic activity and functional specificityQi Chen0Peiguo Yang1College of Life Sciences, Zhejiang University, Hangzhou, Zhejiang, China; Westlake Laboratory of Life Sciences and Biomedicine, Hangzhou, Zhejiang, China; School of Life Sciences, Westlake University, Hangzhou, Zhejiang, ChinaWestlake Laboratory of Life Sciences and Biomedicine, Hangzhou, Zhejiang, China; School of Life Sciences, Westlake University, Hangzhou, Zhejiang, China; Corresponding authorSummary: Tripartite motif (TRIM) proteins are E3 ligases with modular structures involved in diverse cellular functions. Whether biomolecular condensation broadly regulates TRIM proteins remains unclear. Here, we systematically examine 75 TRIMs in mammalian cells and find that the majority can form condensates, either individually or together with other TRIMs. The coiled-coil (CC) domain is essential for TRIM condensation, and disease-related single-nucleotide polymorphisms (SNPs) in this domain impair condensate formation. We show that condensation modulates E3 ligase activity of several TRIMs in a context-dependent manner using cellular auto-ubiquitination and biotinylated-ubiquitination assays. Proteomic analysis reveals that proteins within 21 TRIM condensates are linked to key cellular pathways and diseases. Functional screening uncovers that certain TRIM condensates regulate centriolar satellite organization, cilia assembly, and microtubule stabilization. Our work provides foundational insights into the condensation and functions of TRIM family proteins and establishes a framework for studying co-condensation specificity among highly homologous proteins.http://www.sciencedirect.com/science/article/pii/S2211124725005376CP: Molecular biology
spellingShingle Qi Chen
Peiguo Yang
Coiled-coil-domain-mediated TRIM E3 protein condensation modulates enzymatic activity and functional specificity
Cell Reports
CP: Molecular biology
title Coiled-coil-domain-mediated TRIM E3 protein condensation modulates enzymatic activity and functional specificity
title_full Coiled-coil-domain-mediated TRIM E3 protein condensation modulates enzymatic activity and functional specificity
title_fullStr Coiled-coil-domain-mediated TRIM E3 protein condensation modulates enzymatic activity and functional specificity
title_full_unstemmed Coiled-coil-domain-mediated TRIM E3 protein condensation modulates enzymatic activity and functional specificity
title_short Coiled-coil-domain-mediated TRIM E3 protein condensation modulates enzymatic activity and functional specificity
title_sort coiled coil domain mediated trim e3 protein condensation modulates enzymatic activity and functional specificity
topic CP: Molecular biology
url http://www.sciencedirect.com/science/article/pii/S2211124725005376
work_keys_str_mv AT qichen coiledcoildomainmediatedtrime3proteincondensationmodulatesenzymaticactivityandfunctionalspecificity
AT peiguoyang coiledcoildomainmediatedtrime3proteincondensationmodulatesenzymaticactivityandfunctionalspecificity