A high-resolution haplotype collection uncovers somatic hybridization, recombination and intercontinental movement in oat crown rust.

The population structure and evolution of basidiomycetes like rust fungi are influenced by complex reproductive cycles and dikaryotic life stages where two independent nuclear haplotypes are present in the cell. The ability to alternate between asexual (clonal) and sexual reproduction increases the...

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Main Authors: Eva C Henningsen, David Lewis, Eric S Nazareno, Hayley Mangelson, Monica Sanchez, Kyle Langford, Yung-Fen Huang, Brian J Steffenson, Brendan Boesen, Shahryar F Kianian, Ivan Liachko, Eric Stone, Peter N Dodds, Jana Sperschneider, Melania Figueroa
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2024-11-01
Series:PLoS Genetics
Online Access:https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1011493&type=printable
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author Eva C Henningsen
David Lewis
Eric S Nazareno
Hayley Mangelson
Monica Sanchez
Kyle Langford
Yung-Fen Huang
Brian J Steffenson
Brendan Boesen
Shahryar F Kianian
Ivan Liachko
Eric Stone
Peter N Dodds
Jana Sperschneider
Melania Figueroa
author_facet Eva C Henningsen
David Lewis
Eric S Nazareno
Hayley Mangelson
Monica Sanchez
Kyle Langford
Yung-Fen Huang
Brian J Steffenson
Brendan Boesen
Shahryar F Kianian
Ivan Liachko
Eric Stone
Peter N Dodds
Jana Sperschneider
Melania Figueroa
author_sort Eva C Henningsen
collection DOAJ
description The population structure and evolution of basidiomycetes like rust fungi are influenced by complex reproductive cycles and dikaryotic life stages where two independent nuclear haplotypes are present in the cell. The ability to alternate between asexual (clonal) and sexual reproduction increases the evolutionary capacity in these species. Furthermore, exchange of intact nuclei (somatic hybridization) in rust fungi can allow for rapid generation of genetic variability outside of the sexual cycle. Puccinia coronata f. sp. avenae (Pca), the causal agent of oat crown rust, is a pathogen of global economic importance that is difficult to control due to rapid breakdown of host genetic resistance. The contribution of sexuality, clonality, and migration to virulence evolution varies across Pca populations. As such, the Pca pathosystem is ideal to address the role of mating type, recombination, mutation, and somatic hybridization in host adaptation. We expanded the existing resources for USA and South African populations by generating whole genome sequencing data of Taiwanese and Australian isolates. An atlas of 30 chromosome-level, fully-phased nuclear haplotypes from six USA isolates and nine Australian isolates was created to capture the genomic composition of key Pca lineages. At the haplotype level, we confirmed previous reports of genetic recombination in the USA population and additionally detected either sexual or cryptic recombination between Australian isolates, contrasting previous evaluations that suggested Pca populations in Australia to be purely clonal. We also identified somatic hybridization events in Pca that are not only associated with significant changes in fitness but also imply intercontinental migration of haplotypes, which provides further impetus for molecular monitoring of rust pathogen populations on a global scale.
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spelling doaj-art-bee27104d16f445c9c9456324f4ff0d32025-08-20T02:36:41ZengPublic Library of Science (PLoS)PLoS Genetics1553-73901553-74042024-11-012011e101149310.1371/journal.pgen.1011493A high-resolution haplotype collection uncovers somatic hybridization, recombination and intercontinental movement in oat crown rust.Eva C HenningsenDavid LewisEric S NazarenoHayley MangelsonMonica SanchezKyle LangfordYung-Fen HuangBrian J SteffensonBrendan BoesenShahryar F KianianIvan LiachkoEric StonePeter N DoddsJana SperschneiderMelania FigueroaThe population structure and evolution of basidiomycetes like rust fungi are influenced by complex reproductive cycles and dikaryotic life stages where two independent nuclear haplotypes are present in the cell. The ability to alternate between asexual (clonal) and sexual reproduction increases the evolutionary capacity in these species. Furthermore, exchange of intact nuclei (somatic hybridization) in rust fungi can allow for rapid generation of genetic variability outside of the sexual cycle. Puccinia coronata f. sp. avenae (Pca), the causal agent of oat crown rust, is a pathogen of global economic importance that is difficult to control due to rapid breakdown of host genetic resistance. The contribution of sexuality, clonality, and migration to virulence evolution varies across Pca populations. As such, the Pca pathosystem is ideal to address the role of mating type, recombination, mutation, and somatic hybridization in host adaptation. We expanded the existing resources for USA and South African populations by generating whole genome sequencing data of Taiwanese and Australian isolates. An atlas of 30 chromosome-level, fully-phased nuclear haplotypes from six USA isolates and nine Australian isolates was created to capture the genomic composition of key Pca lineages. At the haplotype level, we confirmed previous reports of genetic recombination in the USA population and additionally detected either sexual or cryptic recombination between Australian isolates, contrasting previous evaluations that suggested Pca populations in Australia to be purely clonal. We also identified somatic hybridization events in Pca that are not only associated with significant changes in fitness but also imply intercontinental migration of haplotypes, which provides further impetus for molecular monitoring of rust pathogen populations on a global scale.https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1011493&type=printable
spellingShingle Eva C Henningsen
David Lewis
Eric S Nazareno
Hayley Mangelson
Monica Sanchez
Kyle Langford
Yung-Fen Huang
Brian J Steffenson
Brendan Boesen
Shahryar F Kianian
Ivan Liachko
Eric Stone
Peter N Dodds
Jana Sperschneider
Melania Figueroa
A high-resolution haplotype collection uncovers somatic hybridization, recombination and intercontinental movement in oat crown rust.
PLoS Genetics
title A high-resolution haplotype collection uncovers somatic hybridization, recombination and intercontinental movement in oat crown rust.
title_full A high-resolution haplotype collection uncovers somatic hybridization, recombination and intercontinental movement in oat crown rust.
title_fullStr A high-resolution haplotype collection uncovers somatic hybridization, recombination and intercontinental movement in oat crown rust.
title_full_unstemmed A high-resolution haplotype collection uncovers somatic hybridization, recombination and intercontinental movement in oat crown rust.
title_short A high-resolution haplotype collection uncovers somatic hybridization, recombination and intercontinental movement in oat crown rust.
title_sort high resolution haplotype collection uncovers somatic hybridization recombination and intercontinental movement in oat crown rust
url https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1011493&type=printable
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