The RNA export factor, Nxt1, is required for tissue specific transcriptional regulation.

The highly conserved, Nxf/Nxt (TAP/p15) RNA nuclear export pathway is important for export of most mRNAs from the nucleus, by interacting with mRNAs and promoting their passage through nuclear pores. Nxt1 is essential for viability; using a partial loss of function allele, we reveal a role for this...

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Main Authors: Simona Caporilli, Yachuan Yu, Jianqiao Jiang, Helen White-Cooper
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2013-06-01
Series:PLoS Genetics
Online Access:https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1003526&type=printable
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author Simona Caporilli
Yachuan Yu
Jianqiao Jiang
Helen White-Cooper
author_facet Simona Caporilli
Yachuan Yu
Jianqiao Jiang
Helen White-Cooper
author_sort Simona Caporilli
collection DOAJ
description The highly conserved, Nxf/Nxt (TAP/p15) RNA nuclear export pathway is important for export of most mRNAs from the nucleus, by interacting with mRNAs and promoting their passage through nuclear pores. Nxt1 is essential for viability; using a partial loss of function allele, we reveal a role for this gene in tissue specific transcription. We show that many Drosophila melanogaster testis-specific mRNAs require Nxt1 for their accumulation. The transcripts that require Nxt1 also depend on a testis-specific transcription complex, tMAC. We show that loss of Nxt1 leads to reduced transcription of tMAC targets. A reporter transcript from a tMAC-dependent promoter is under-expressed in Nxt1 mutants, however the same transcript accumulates in mutants if driven by a tMAC-independent promoter. Thus, in Drosophila primary spermatocytes, the transcription factor used to activate expression of a transcript, rather than the RNA sequence itself or the core transcription machinery, determines whether this expression requires Nxt1. We additionally find that transcripts from intron-less genes are more sensitive to loss of Nxt1 function than those from intron-containing genes and propose a mechanism in which transcript processing feeds back to increase activity of a tissue specific transcription complex.
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spelling doaj-art-bebd35eaf0ab43d1bde539c009ec561f2025-08-20T02:33:43ZengPublic Library of Science (PLoS)PLoS Genetics1553-73901553-74042013-06-0196e100352610.1371/journal.pgen.1003526The RNA export factor, Nxt1, is required for tissue specific transcriptional regulation.Simona CaporilliYachuan YuJianqiao JiangHelen White-CooperThe highly conserved, Nxf/Nxt (TAP/p15) RNA nuclear export pathway is important for export of most mRNAs from the nucleus, by interacting with mRNAs and promoting their passage through nuclear pores. Nxt1 is essential for viability; using a partial loss of function allele, we reveal a role for this gene in tissue specific transcription. We show that many Drosophila melanogaster testis-specific mRNAs require Nxt1 for their accumulation. The transcripts that require Nxt1 also depend on a testis-specific transcription complex, tMAC. We show that loss of Nxt1 leads to reduced transcription of tMAC targets. A reporter transcript from a tMAC-dependent promoter is under-expressed in Nxt1 mutants, however the same transcript accumulates in mutants if driven by a tMAC-independent promoter. Thus, in Drosophila primary spermatocytes, the transcription factor used to activate expression of a transcript, rather than the RNA sequence itself or the core transcription machinery, determines whether this expression requires Nxt1. We additionally find that transcripts from intron-less genes are more sensitive to loss of Nxt1 function than those from intron-containing genes and propose a mechanism in which transcript processing feeds back to increase activity of a tissue specific transcription complex.https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1003526&type=printable
spellingShingle Simona Caporilli
Yachuan Yu
Jianqiao Jiang
Helen White-Cooper
The RNA export factor, Nxt1, is required for tissue specific transcriptional regulation.
PLoS Genetics
title The RNA export factor, Nxt1, is required for tissue specific transcriptional regulation.
title_full The RNA export factor, Nxt1, is required for tissue specific transcriptional regulation.
title_fullStr The RNA export factor, Nxt1, is required for tissue specific transcriptional regulation.
title_full_unstemmed The RNA export factor, Nxt1, is required for tissue specific transcriptional regulation.
title_short The RNA export factor, Nxt1, is required for tissue specific transcriptional regulation.
title_sort rna export factor nxt1 is required for tissue specific transcriptional regulation
url https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1003526&type=printable
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