Virus-modified paraspeckle-like condensates are hubs for viral RNA processing and their formation drives genomic instability

Abstract The nucleus is a highly organised yet dynamic environment containing distinct membraneless nuclear bodies. This spatial separation enables a subset of components to be concentrated within biomolecular condensates, allowing efficient and discrete processes to occur which regulate cellular fu...

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Main Authors: Katherine L. Harper, Elena M. Harrington, Connor Hayward, Chinedu A. Anene, Wiyada Wongwiwat, Robert E. White, Adrian Whitehouse
Format: Article
Language:English
Published: Nature Portfolio 2024-11-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-024-54592-5
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author Katherine L. Harper
Elena M. Harrington
Connor Hayward
Chinedu A. Anene
Wiyada Wongwiwat
Robert E. White
Adrian Whitehouse
author_facet Katherine L. Harper
Elena M. Harrington
Connor Hayward
Chinedu A. Anene
Wiyada Wongwiwat
Robert E. White
Adrian Whitehouse
author_sort Katherine L. Harper
collection DOAJ
description Abstract The nucleus is a highly organised yet dynamic environment containing distinct membraneless nuclear bodies. This spatial separation enables a subset of components to be concentrated within biomolecular condensates, allowing efficient and discrete processes to occur which regulate cellular function. One such nuclear body, paraspeckles, are comprised of multiple paraspeckle proteins (PSPs) built around the architectural RNA, NEAT1_2. Paraspeckle function is yet to be fully elucidated but has been implicated in a variety of developmental and disease scenarios. We demonstrate that Kaposi’s sarcoma-associated herpesvirus (KSHV) drives formation of structurally distinct paraspeckles with a dramatically increased size and altered protein composition that are required for productive lytic replication. We highlight these virus-modified paraspeckles form adjacent to virus replication centres, potentially functioning as RNA processing hubs for viral transcripts during infection. Notably, we reveal that PSP sequestration into virus-modified paraspeckles result in increased genome instability during both KSHV and Epstein Barr virus (EBV) infection, implicating their formation in virus-mediated tumourigenesis.
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spelling doaj-art-bbc07e1451e846af9302412cbd768d192025-08-20T02:08:20ZengNature PortfolioNature Communications2041-17232024-11-0115112510.1038/s41467-024-54592-5Virus-modified paraspeckle-like condensates are hubs for viral RNA processing and their formation drives genomic instabilityKatherine L. Harper0Elena M. Harrington1Connor Hayward2Chinedu A. Anene3Wiyada Wongwiwat4Robert E. White5Adrian Whitehouse6School of Molecular and Cellular Biology and Astbury Centre for Structural Molecular Biology, University of LeedsSchool of Molecular and Cellular Biology and Astbury Centre for Structural Molecular Biology, University of LeedsSchool of Molecular and Cellular Biology and Astbury Centre for Structural Molecular Biology, University of LeedsCentre for Biomedical Science Research, School of Health, Leeds Beckett UniversityDepartment of Infectious Disease, Imperial College London, South Kensington CampusDepartment of Infectious Disease, Imperial College London, South Kensington CampusSchool of Molecular and Cellular Biology and Astbury Centre for Structural Molecular Biology, University of LeedsAbstract The nucleus is a highly organised yet dynamic environment containing distinct membraneless nuclear bodies. This spatial separation enables a subset of components to be concentrated within biomolecular condensates, allowing efficient and discrete processes to occur which regulate cellular function. One such nuclear body, paraspeckles, are comprised of multiple paraspeckle proteins (PSPs) built around the architectural RNA, NEAT1_2. Paraspeckle function is yet to be fully elucidated but has been implicated in a variety of developmental and disease scenarios. We demonstrate that Kaposi’s sarcoma-associated herpesvirus (KSHV) drives formation of structurally distinct paraspeckles with a dramatically increased size and altered protein composition that are required for productive lytic replication. We highlight these virus-modified paraspeckles form adjacent to virus replication centres, potentially functioning as RNA processing hubs for viral transcripts during infection. Notably, we reveal that PSP sequestration into virus-modified paraspeckles result in increased genome instability during both KSHV and Epstein Barr virus (EBV) infection, implicating their formation in virus-mediated tumourigenesis.https://doi.org/10.1038/s41467-024-54592-5
spellingShingle Katherine L. Harper
Elena M. Harrington
Connor Hayward
Chinedu A. Anene
Wiyada Wongwiwat
Robert E. White
Adrian Whitehouse
Virus-modified paraspeckle-like condensates are hubs for viral RNA processing and their formation drives genomic instability
Nature Communications
title Virus-modified paraspeckle-like condensates are hubs for viral RNA processing and their formation drives genomic instability
title_full Virus-modified paraspeckle-like condensates are hubs for viral RNA processing and their formation drives genomic instability
title_fullStr Virus-modified paraspeckle-like condensates are hubs for viral RNA processing and their formation drives genomic instability
title_full_unstemmed Virus-modified paraspeckle-like condensates are hubs for viral RNA processing and their formation drives genomic instability
title_short Virus-modified paraspeckle-like condensates are hubs for viral RNA processing and their formation drives genomic instability
title_sort virus modified paraspeckle like condensates are hubs for viral rna processing and their formation drives genomic instability
url https://doi.org/10.1038/s41467-024-54592-5
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