Epigenomic and 3D genomic mapping reveals developmental dynamics and subgenomic asymmetry of transcriptional regulatory architecture in allotetraploid cotton

Abstract Although epigenetic modification has long been recognized as a vital force influencing gene regulation in plants, the dynamics of chromatin structure implicated in the intertwined transcriptional regulation of duplicated genes in polyploids have yet to be understood. Here, we document the d...

Full description

Saved in:
Bibliographic Details
Main Authors: Xianhui Huang, Yuejin Wang, Sainan Zhang, Liuling Pei, Jiaqi You, Yuexuan Long, Jianying Li, Xianlong Zhang, Longfu Zhu, Maojun Wang
Format: Article
Language:English
Published: Nature Portfolio 2024-12-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-024-55309-4
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1850102928806772736
author Xianhui Huang
Yuejin Wang
Sainan Zhang
Liuling Pei
Jiaqi You
Yuexuan Long
Jianying Li
Xianlong Zhang
Longfu Zhu
Maojun Wang
author_facet Xianhui Huang
Yuejin Wang
Sainan Zhang
Liuling Pei
Jiaqi You
Yuexuan Long
Jianying Li
Xianlong Zhang
Longfu Zhu
Maojun Wang
author_sort Xianhui Huang
collection DOAJ
description Abstract Although epigenetic modification has long been recognized as a vital force influencing gene regulation in plants, the dynamics of chromatin structure implicated in the intertwined transcriptional regulation of duplicated genes in polyploids have yet to be understood. Here, we document the dynamic organization of chromatin structure in two subgenomes of allotetraploid cotton (Gossypium hirsutum) by generating 3D genomic, epigenomic and transcriptomic datasets from 12 major tissues/developmental stages covering the life cycle. We systematically identify a subset of genes that are closely associated with specific tissue functions. Interestingly, these genes exhibit not only higher tissue specificity but also a more pronounced homoeologous bias. We comprehensively elucidate the intricate process of subgenomic collaboration and divergence across various tissues. A comparison among subgenomes in the 12 tissues reveals widespread differences in the reorganization of 3D genome structures, with the Dt subgenome exhibiting a higher extent of dynamic chromatin status than the At subgenome. Moreover, we construct a comprehensive atlas of putative functional genome elements and discover that 37 cis-regulatory elements (CREs) have selection signals acquired during domestication and improvement. These data and analyses are publicly available to the research community through a web portal. In summary, this study provides abundant resources and depicts the regulatory architecture of the genome, which thereby facilitates the understanding of biological processes and guides cotton breeding.
format Article
id doaj-art-b986cef600fc4005ae3f03d10dda8427
institution DOAJ
issn 2041-1723
language English
publishDate 2024-12-01
publisher Nature Portfolio
record_format Article
series Nature Communications
spelling doaj-art-b986cef600fc4005ae3f03d10dda84272025-08-20T02:39:38ZengNature PortfolioNature Communications2041-17232024-12-0115112010.1038/s41467-024-55309-4Epigenomic and 3D genomic mapping reveals developmental dynamics and subgenomic asymmetry of transcriptional regulatory architecture in allotetraploid cottonXianhui Huang0Yuejin Wang1Sainan Zhang2Liuling Pei3Jiaqi You4Yuexuan Long5Jianying Li6Xianlong Zhang7Longfu Zhu8Maojun Wang9National Key Laboratory of Crop Genetic Improvement, Hubei Hongshan Laboratory, Huazhong Agricultural UniversityNational Key Laboratory of Crop Genetic Improvement, Hubei Hongshan Laboratory, Huazhong Agricultural UniversityNational Key Laboratory of Crop Genetic Improvement, Hubei Hongshan Laboratory, Huazhong Agricultural UniversityNational Key Laboratory of Crop Genetic Improvement, Hubei Hongshan Laboratory, Huazhong Agricultural UniversityNational Key Laboratory of Crop Genetic Improvement, Hubei Hongshan Laboratory, Huazhong Agricultural UniversityNational Key Laboratory of Crop Genetic Improvement, Hubei Hongshan Laboratory, Huazhong Agricultural UniversityNational Key Laboratory of Crop Genetic Improvement, Hubei Hongshan Laboratory, Huazhong Agricultural UniversityNational Key Laboratory of Crop Genetic Improvement, Hubei Hongshan Laboratory, Huazhong Agricultural UniversityNational Key Laboratory of Crop Genetic Improvement, Hubei Hongshan Laboratory, Huazhong Agricultural UniversityNational Key Laboratory of Crop Genetic Improvement, Hubei Hongshan Laboratory, Huazhong Agricultural UniversityAbstract Although epigenetic modification has long been recognized as a vital force influencing gene regulation in plants, the dynamics of chromatin structure implicated in the intertwined transcriptional regulation of duplicated genes in polyploids have yet to be understood. Here, we document the dynamic organization of chromatin structure in two subgenomes of allotetraploid cotton (Gossypium hirsutum) by generating 3D genomic, epigenomic and transcriptomic datasets from 12 major tissues/developmental stages covering the life cycle. We systematically identify a subset of genes that are closely associated with specific tissue functions. Interestingly, these genes exhibit not only higher tissue specificity but also a more pronounced homoeologous bias. We comprehensively elucidate the intricate process of subgenomic collaboration and divergence across various tissues. A comparison among subgenomes in the 12 tissues reveals widespread differences in the reorganization of 3D genome structures, with the Dt subgenome exhibiting a higher extent of dynamic chromatin status than the At subgenome. Moreover, we construct a comprehensive atlas of putative functional genome elements and discover that 37 cis-regulatory elements (CREs) have selection signals acquired during domestication and improvement. These data and analyses are publicly available to the research community through a web portal. In summary, this study provides abundant resources and depicts the regulatory architecture of the genome, which thereby facilitates the understanding of biological processes and guides cotton breeding.https://doi.org/10.1038/s41467-024-55309-4
spellingShingle Xianhui Huang
Yuejin Wang
Sainan Zhang
Liuling Pei
Jiaqi You
Yuexuan Long
Jianying Li
Xianlong Zhang
Longfu Zhu
Maojun Wang
Epigenomic and 3D genomic mapping reveals developmental dynamics and subgenomic asymmetry of transcriptional regulatory architecture in allotetraploid cotton
Nature Communications
title Epigenomic and 3D genomic mapping reveals developmental dynamics and subgenomic asymmetry of transcriptional regulatory architecture in allotetraploid cotton
title_full Epigenomic and 3D genomic mapping reveals developmental dynamics and subgenomic asymmetry of transcriptional regulatory architecture in allotetraploid cotton
title_fullStr Epigenomic and 3D genomic mapping reveals developmental dynamics and subgenomic asymmetry of transcriptional regulatory architecture in allotetraploid cotton
title_full_unstemmed Epigenomic and 3D genomic mapping reveals developmental dynamics and subgenomic asymmetry of transcriptional regulatory architecture in allotetraploid cotton
title_short Epigenomic and 3D genomic mapping reveals developmental dynamics and subgenomic asymmetry of transcriptional regulatory architecture in allotetraploid cotton
title_sort epigenomic and 3d genomic mapping reveals developmental dynamics and subgenomic asymmetry of transcriptional regulatory architecture in allotetraploid cotton
url https://doi.org/10.1038/s41467-024-55309-4
work_keys_str_mv AT xianhuihuang epigenomicand3dgenomicmappingrevealsdevelopmentaldynamicsandsubgenomicasymmetryoftranscriptionalregulatoryarchitectureinallotetraploidcotton
AT yuejinwang epigenomicand3dgenomicmappingrevealsdevelopmentaldynamicsandsubgenomicasymmetryoftranscriptionalregulatoryarchitectureinallotetraploidcotton
AT sainanzhang epigenomicand3dgenomicmappingrevealsdevelopmentaldynamicsandsubgenomicasymmetryoftranscriptionalregulatoryarchitectureinallotetraploidcotton
AT liulingpei epigenomicand3dgenomicmappingrevealsdevelopmentaldynamicsandsubgenomicasymmetryoftranscriptionalregulatoryarchitectureinallotetraploidcotton
AT jiaqiyou epigenomicand3dgenomicmappingrevealsdevelopmentaldynamicsandsubgenomicasymmetryoftranscriptionalregulatoryarchitectureinallotetraploidcotton
AT yuexuanlong epigenomicand3dgenomicmappingrevealsdevelopmentaldynamicsandsubgenomicasymmetryoftranscriptionalregulatoryarchitectureinallotetraploidcotton
AT jianyingli epigenomicand3dgenomicmappingrevealsdevelopmentaldynamicsandsubgenomicasymmetryoftranscriptionalregulatoryarchitectureinallotetraploidcotton
AT xianlongzhang epigenomicand3dgenomicmappingrevealsdevelopmentaldynamicsandsubgenomicasymmetryoftranscriptionalregulatoryarchitectureinallotetraploidcotton
AT longfuzhu epigenomicand3dgenomicmappingrevealsdevelopmentaldynamicsandsubgenomicasymmetryoftranscriptionalregulatoryarchitectureinallotetraploidcotton
AT maojunwang epigenomicand3dgenomicmappingrevealsdevelopmentaldynamicsandsubgenomicasymmetryoftranscriptionalregulatoryarchitectureinallotetraploidcotton