Transcription regulation of African swine fever virus: dual role of M1249L

Abstract African swine fever virus (ASFV), which poses significant risks to the global economy, encodes a unique host-independent transcription system. This system comprises an eight-subunit RNA polymerase (vRNAP), temporally expressed transcription factors and transcript associated proteins, facili...

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Main Authors: Dongming Zhao, Nan Wang, Xiaoying Feng, Zhenjiang Zhang, Kongen Xu, Tao Zheng, Yunge Yang, Xuemei Li, Xianjin Ou, Rui Zhao, Zihe Rao, Zhigao Bu, Yutao Chen, Xiangxi Wang
Format: Article
Language:English
Published: Nature Portfolio 2024-11-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-024-54461-1
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author Dongming Zhao
Nan Wang
Xiaoying Feng
Zhenjiang Zhang
Kongen Xu
Tao Zheng
Yunge Yang
Xuemei Li
Xianjin Ou
Rui Zhao
Zihe Rao
Zhigao Bu
Yutao Chen
Xiangxi Wang
author_facet Dongming Zhao
Nan Wang
Xiaoying Feng
Zhenjiang Zhang
Kongen Xu
Tao Zheng
Yunge Yang
Xuemei Li
Xianjin Ou
Rui Zhao
Zihe Rao
Zhigao Bu
Yutao Chen
Xiangxi Wang
author_sort Dongming Zhao
collection DOAJ
description Abstract African swine fever virus (ASFV), which poses significant risks to the global economy, encodes a unique host-independent transcription system. This system comprises an eight-subunit RNA polymerase (vRNAP), temporally expressed transcription factors and transcript associated proteins, facilitating cross-species transmission via intermediate host. The protein composition of the virion and the presence of transcription factors in virus genome suggest existence of distinct transcription systems during viral infection. However, the precise mechanisms of transcription regulation remain elusive. Through analyses of dynamic transcriptome, vRNAP-associated components and cell-based assay, the critical role of M1249L in viral transcription regulation has been highlighted. Atomic-resolution structures of vRNAP-M1249L supercomplex, exhibiting a variety of conformations, have uncovered the dual functions of M1249L. During early transcription, M1249L could serve as multiple temporary transcription factors with C-terminal domain acting as a switcher for activation/inactivation, while during late transcription it aids in the packaging of the transcription machinery. The structural and functional characteristics of M1249L underscore its vital roles in ASFV transcription, packaging, and capsid assembly, presenting novel opportunities for therapeutic intervention.
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issn 2041-1723
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publishDate 2024-11-01
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spelling doaj-art-b897bb29a9894c7e9c58c1ce0ba292d72024-11-24T12:33:49ZengNature PortfolioNature Communications2041-17232024-11-0115111410.1038/s41467-024-54461-1Transcription regulation of African swine fever virus: dual role of M1249LDongming Zhao0Nan Wang1Xiaoying Feng2Zhenjiang Zhang3Kongen Xu4Tao Zheng5Yunge Yang6Xuemei Li7Xianjin Ou8Rui Zhao9Zihe Rao10Zhigao Bu11Yutao Chen12Xiangxi Wang13State Key Laboratory for Animal Disease Control and Prevention, National African Swine Fever Para-reference Laboratory, National High Containment Facilities for Animal Diseases Control and Prevention, Harbin Veterinary Research Institute, Chinese Academy of Agricultural SciencesNational Laboratory of Biomacromolecules, Key Laboratory of Infection and Immunity, Institute of Biophysics, Chinese Academy of SciencesNational Laboratory of Biomacromolecules, Key Laboratory of Infection and Immunity, Institute of Biophysics, Chinese Academy of SciencesState Key Laboratory for Animal Disease Control and Prevention, National African Swine Fever Para-reference Laboratory, National High Containment Facilities for Animal Diseases Control and Prevention, Harbin Veterinary Research Institute, Chinese Academy of Agricultural SciencesNational Laboratory of Biomacromolecules, Key Laboratory of Infection and Immunity, Institute of Biophysics, Chinese Academy of SciencesNational Laboratory of Biomacromolecules, Key Laboratory of Infection and Immunity, Institute of Biophysics, Chinese Academy of SciencesNational Laboratory of Biomacromolecules, Key Laboratory of Infection and Immunity, Institute of Biophysics, Chinese Academy of SciencesNational Laboratory of Biomacromolecules, Key Laboratory of Infection and Immunity, Institute of Biophysics, Chinese Academy of SciencesNational Laboratory of Biomacromolecules, Key Laboratory of Infection and Immunity, Institute of Biophysics, Chinese Academy of SciencesState Key Laboratory for Animal Disease Control and Prevention, National African Swine Fever Para-reference Laboratory, National High Containment Facilities for Animal Diseases Control and Prevention, Harbin Veterinary Research Institute, Chinese Academy of Agricultural SciencesNational Laboratory of Biomacromolecules, Key Laboratory of Infection and Immunity, Institute of Biophysics, Chinese Academy of SciencesState Key Laboratory for Animal Disease Control and Prevention, National African Swine Fever Para-reference Laboratory, National High Containment Facilities for Animal Diseases Control and Prevention, Harbin Veterinary Research Institute, Chinese Academy of Agricultural SciencesNational Laboratory of Biomacromolecules, Key Laboratory of Infection and Immunity, Institute of Biophysics, Chinese Academy of SciencesNational Laboratory of Biomacromolecules, Key Laboratory of Infection and Immunity, Institute of Biophysics, Chinese Academy of SciencesAbstract African swine fever virus (ASFV), which poses significant risks to the global economy, encodes a unique host-independent transcription system. This system comprises an eight-subunit RNA polymerase (vRNAP), temporally expressed transcription factors and transcript associated proteins, facilitating cross-species transmission via intermediate host. The protein composition of the virion and the presence of transcription factors in virus genome suggest existence of distinct transcription systems during viral infection. However, the precise mechanisms of transcription regulation remain elusive. Through analyses of dynamic transcriptome, vRNAP-associated components and cell-based assay, the critical role of M1249L in viral transcription regulation has been highlighted. Atomic-resolution structures of vRNAP-M1249L supercomplex, exhibiting a variety of conformations, have uncovered the dual functions of M1249L. During early transcription, M1249L could serve as multiple temporary transcription factors with C-terminal domain acting as a switcher for activation/inactivation, while during late transcription it aids in the packaging of the transcription machinery. The structural and functional characteristics of M1249L underscore its vital roles in ASFV transcription, packaging, and capsid assembly, presenting novel opportunities for therapeutic intervention.https://doi.org/10.1038/s41467-024-54461-1
spellingShingle Dongming Zhao
Nan Wang
Xiaoying Feng
Zhenjiang Zhang
Kongen Xu
Tao Zheng
Yunge Yang
Xuemei Li
Xianjin Ou
Rui Zhao
Zihe Rao
Zhigao Bu
Yutao Chen
Xiangxi Wang
Transcription regulation of African swine fever virus: dual role of M1249L
Nature Communications
title Transcription regulation of African swine fever virus: dual role of M1249L
title_full Transcription regulation of African swine fever virus: dual role of M1249L
title_fullStr Transcription regulation of African swine fever virus: dual role of M1249L
title_full_unstemmed Transcription regulation of African swine fever virus: dual role of M1249L
title_short Transcription regulation of African swine fever virus: dual role of M1249L
title_sort transcription regulation of african swine fever virus dual role of m1249l
url https://doi.org/10.1038/s41467-024-54461-1
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