hnRNP I inhibits Notch signaling and regulates intestinal epithelial homeostasis in the zebrafish.

Regulated intestinal stem cell proliferation and differentiation are required for normal intestinal homeostasis and repair after injury. The Notch signaling pathway plays fundamental roles in the intestinal epithelium. Despite the fact that Notch signaling maintains intestinal stem cells in a prolif...

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Main Authors: Jing Yang, Chin Yee Chan, Bo Jiang, Xueyuan Yu, Guo-Zhang Zhu, Yiping Chen, John Barnard, Wenyan Mei
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2009-02-01
Series:PLoS Genetics
Online Access:https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1000363&type=printable
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author Jing Yang
Chin Yee Chan
Bo Jiang
Xueyuan Yu
Guo-Zhang Zhu
Yiping Chen
John Barnard
Wenyan Mei
author_facet Jing Yang
Chin Yee Chan
Bo Jiang
Xueyuan Yu
Guo-Zhang Zhu
Yiping Chen
John Barnard
Wenyan Mei
author_sort Jing Yang
collection DOAJ
description Regulated intestinal stem cell proliferation and differentiation are required for normal intestinal homeostasis and repair after injury. The Notch signaling pathway plays fundamental roles in the intestinal epithelium. Despite the fact that Notch signaling maintains intestinal stem cells in a proliferative state and promotes absorptive cell differentiation in most species, it remains largely unclear how Notch signaling itself is precisely controlled during intestinal homeostasis. We characterized the intestinal phenotypes of brom bones, a zebrafish mutant carrying a nonsense mutation in hnRNP I. We found that the brom bones mutant displays a number of intestinal defects, including compromised secretory goblet cell differentiation, hyperproliferation, and enhanced apoptosis. These phenotypes are accompanied by a markedly elevated Notch signaling activity in the intestinal epithelium. When overexpressed, hnRNP I destabilizes the Notch intracellular domain (NICD) and inhibits Notch signaling. This activity of hnRNP I is conserved from zebrafish to human. In addition, our biochemistry experiments demonstrate that the effect of hnRNP I on NICD turnover requires the C-terminal portion of the RAM domain of NICD. Our results demonstrate that hnRNP I is an evolutionarily conserved Notch inhibitor and plays an essential role in intestinal homeostasis.
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issn 1553-7390
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language English
publishDate 2009-02-01
publisher Public Library of Science (PLoS)
record_format Article
series PLoS Genetics
spelling doaj-art-b38123731223480ba0d1c86ccce346c72025-08-20T02:38:29ZengPublic Library of Science (PLoS)PLoS Genetics1553-73901553-74042009-02-0152e100036310.1371/journal.pgen.1000363hnRNP I inhibits Notch signaling and regulates intestinal epithelial homeostasis in the zebrafish.Jing YangChin Yee ChanBo JiangXueyuan YuGuo-Zhang ZhuYiping ChenJohn BarnardWenyan MeiRegulated intestinal stem cell proliferation and differentiation are required for normal intestinal homeostasis and repair after injury. The Notch signaling pathway plays fundamental roles in the intestinal epithelium. Despite the fact that Notch signaling maintains intestinal stem cells in a proliferative state and promotes absorptive cell differentiation in most species, it remains largely unclear how Notch signaling itself is precisely controlled during intestinal homeostasis. We characterized the intestinal phenotypes of brom bones, a zebrafish mutant carrying a nonsense mutation in hnRNP I. We found that the brom bones mutant displays a number of intestinal defects, including compromised secretory goblet cell differentiation, hyperproliferation, and enhanced apoptosis. These phenotypes are accompanied by a markedly elevated Notch signaling activity in the intestinal epithelium. When overexpressed, hnRNP I destabilizes the Notch intracellular domain (NICD) and inhibits Notch signaling. This activity of hnRNP I is conserved from zebrafish to human. In addition, our biochemistry experiments demonstrate that the effect of hnRNP I on NICD turnover requires the C-terminal portion of the RAM domain of NICD. Our results demonstrate that hnRNP I is an evolutionarily conserved Notch inhibitor and plays an essential role in intestinal homeostasis.https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1000363&type=printable
spellingShingle Jing Yang
Chin Yee Chan
Bo Jiang
Xueyuan Yu
Guo-Zhang Zhu
Yiping Chen
John Barnard
Wenyan Mei
hnRNP I inhibits Notch signaling and regulates intestinal epithelial homeostasis in the zebrafish.
PLoS Genetics
title hnRNP I inhibits Notch signaling and regulates intestinal epithelial homeostasis in the zebrafish.
title_full hnRNP I inhibits Notch signaling and regulates intestinal epithelial homeostasis in the zebrafish.
title_fullStr hnRNP I inhibits Notch signaling and regulates intestinal epithelial homeostasis in the zebrafish.
title_full_unstemmed hnRNP I inhibits Notch signaling and regulates intestinal epithelial homeostasis in the zebrafish.
title_short hnRNP I inhibits Notch signaling and regulates intestinal epithelial homeostasis in the zebrafish.
title_sort hnrnp i inhibits notch signaling and regulates intestinal epithelial homeostasis in the zebrafish
url https://journals.plos.org/plosgenetics/article/file?id=10.1371/journal.pgen.1000363&type=printable
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