Genome-wide identification and analysis of the FLA gene family in maize (Zea mays L.) and its expression in response to abiotic stresses

Abstract Background Fasciclin-like arabinogalactan proteins (FLAs) are a class of plant-specific proteins that contribute to various aspects of plant growth, development, and adaptation. However, these proteins have not yet been identified or characterized in maize. This study conducted bioinformati...

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Main Authors: Yongyan Cao, Xuanxuan Chen, Bo Wei, Tuo Zeng, Hongcheng Wang, Bin Zhu, Xuye Du, Lei Gu
Format: Article
Language:English
Published: BMC 2025-07-01
Series:BMC Plant Biology
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Online Access:https://doi.org/10.1186/s12870-025-06981-1
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author Yongyan Cao
Xuanxuan Chen
Bo Wei
Tuo Zeng
Hongcheng Wang
Bin Zhu
Xuye Du
Lei Gu
author_facet Yongyan Cao
Xuanxuan Chen
Bo Wei
Tuo Zeng
Hongcheng Wang
Bin Zhu
Xuye Du
Lei Gu
author_sort Yongyan Cao
collection DOAJ
description Abstract Background Fasciclin-like arabinogalactan proteins (FLAs) are a class of plant-specific proteins that contribute to various aspects of plant growth, development, and adaptation. However, these proteins have not yet been identified or characterized in maize. This study conducted bioinformatics and expression analyses of the ZmFLA gene family. Results Based on the homologous comparison methods, 27 ZmFLAs were identified in maize. They were unevenly distributed across ten maize chromosomes, forming 14 pairs of fragment replication with no tandem duplication, revealing that segmental duplication might be the main driving force of the family expansion. Ka/ks analysis of duplicated genes indicating ZmFLAs may undergoing both positively selected and purification selection in evolutionary history. Homologous evolutionary analysis classified 109 FLAs from maize (27 ZmFLAs), rice (27 OsFLAs), wheat (34 TaFLAs), and Arabidopsis (21 AtFLAs) into three groups. The gene structure and protein domain of the ZmFLAs in the same subfamily were highly conserved. The promoter regions of many ZmFLA family members comprised light-, plant growth/hormone-, and abiotic stress-responsive elements. RNA-seq data revealed that the expression pattern of ZmFLAs exhibited organizational specificity and could be influenced by heat, cold, drought, UV, and salt stresses. RT-qPCR of ten ZmFLAs (ZmFLA1/4/5/9/11/16/19/23/25/27) showed that they were either positively or negatively regulated by drought and salt stresses. These findings suggested that ZmFLAs might be involved in the maize response to drought and salt stresses. Conclusion Taken together, the comprehensive analysis of ZmFLAs in the whole genome provides a novel perspective on the evolutionary relationship among maize FLA genes and lays a foundation for further analysis of the biological functions of ZmFLAs.
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spelling doaj-art-b0f24bd9c2594f6293167e904451f9132025-08-20T03:42:35ZengBMCBMC Plant Biology1471-22292025-07-0125111510.1186/s12870-025-06981-1Genome-wide identification and analysis of the FLA gene family in maize (Zea mays L.) and its expression in response to abiotic stressesYongyan Cao0Xuanxuan Chen1Bo Wei2Tuo Zeng3Hongcheng Wang4Bin Zhu5Xuye Du6Lei Gu7School of Life Sciences, Guizhou Normal UniversitySchool of Life Sciences, Guizhou Normal UniversitySchool of Life Sciences, Guizhou Normal UniversitySchool of Life Sciences, Guizhou Normal UniversitySchool of Life Sciences, Guizhou Normal UniversitySchool of Life Sciences, Guizhou Normal UniversitySchool of Life Sciences, Guizhou Normal UniversitySchool of Life Sciences, Guizhou Normal UniversityAbstract Background Fasciclin-like arabinogalactan proteins (FLAs) are a class of plant-specific proteins that contribute to various aspects of plant growth, development, and adaptation. However, these proteins have not yet been identified or characterized in maize. This study conducted bioinformatics and expression analyses of the ZmFLA gene family. Results Based on the homologous comparison methods, 27 ZmFLAs were identified in maize. They were unevenly distributed across ten maize chromosomes, forming 14 pairs of fragment replication with no tandem duplication, revealing that segmental duplication might be the main driving force of the family expansion. Ka/ks analysis of duplicated genes indicating ZmFLAs may undergoing both positively selected and purification selection in evolutionary history. Homologous evolutionary analysis classified 109 FLAs from maize (27 ZmFLAs), rice (27 OsFLAs), wheat (34 TaFLAs), and Arabidopsis (21 AtFLAs) into three groups. The gene structure and protein domain of the ZmFLAs in the same subfamily were highly conserved. The promoter regions of many ZmFLA family members comprised light-, plant growth/hormone-, and abiotic stress-responsive elements. RNA-seq data revealed that the expression pattern of ZmFLAs exhibited organizational specificity and could be influenced by heat, cold, drought, UV, and salt stresses. RT-qPCR of ten ZmFLAs (ZmFLA1/4/5/9/11/16/19/23/25/27) showed that they were either positively or negatively regulated by drought and salt stresses. These findings suggested that ZmFLAs might be involved in the maize response to drought and salt stresses. Conclusion Taken together, the comprehensive analysis of ZmFLAs in the whole genome provides a novel perspective on the evolutionary relationship among maize FLA genes and lays a foundation for further analysis of the biological functions of ZmFLAs.https://doi.org/10.1186/s12870-025-06981-1MaizeFLA proteinGenome-wideBioinformatics analysisDrought and salt stresses
spellingShingle Yongyan Cao
Xuanxuan Chen
Bo Wei
Tuo Zeng
Hongcheng Wang
Bin Zhu
Xuye Du
Lei Gu
Genome-wide identification and analysis of the FLA gene family in maize (Zea mays L.) and its expression in response to abiotic stresses
BMC Plant Biology
Maize
FLA protein
Genome-wide
Bioinformatics analysis
Drought and salt stresses
title Genome-wide identification and analysis of the FLA gene family in maize (Zea mays L.) and its expression in response to abiotic stresses
title_full Genome-wide identification and analysis of the FLA gene family in maize (Zea mays L.) and its expression in response to abiotic stresses
title_fullStr Genome-wide identification and analysis of the FLA gene family in maize (Zea mays L.) and its expression in response to abiotic stresses
title_full_unstemmed Genome-wide identification and analysis of the FLA gene family in maize (Zea mays L.) and its expression in response to abiotic stresses
title_short Genome-wide identification and analysis of the FLA gene family in maize (Zea mays L.) and its expression in response to abiotic stresses
title_sort genome wide identification and analysis of the fla gene family in maize zea mays l and its expression in response to abiotic stresses
topic Maize
FLA protein
Genome-wide
Bioinformatics analysis
Drought and salt stresses
url https://doi.org/10.1186/s12870-025-06981-1
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