A stochastic explanation for observed local-to-global foraging states in Caenorhabditis elegans

Abrupt changes in behavior can often be associated with changes in underlying behavioral states. When placed off food, the foraging behavior of C. elegans can be described as a change between an initial local-search behavior characterized by a high rate of reorientations, followed by a global-search...

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Main Authors: Andrew Margolis, Andrew Gordus
Format: Article
Language:English
Published: eLife Sciences Publications Ltd 2025-07-01
Series:eLife
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Online Access:https://elifesciences.org/articles/104972
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author Andrew Margolis
Andrew Gordus
author_facet Andrew Margolis
Andrew Gordus
author_sort Andrew Margolis
collection DOAJ
description Abrupt changes in behavior can often be associated with changes in underlying behavioral states. When placed off food, the foraging behavior of C. elegans can be described as a change between an initial local-search behavior characterized by a high rate of reorientations, followed by a global-search behavior characterized by sparse reorientations. This is commonly observed in individual worms, but when numerous worms are characterized, only about half appear to exhibit this behavior. We propose an alternative model that predicts both abrupt and continuous changes to reorientation that do not rely on behavioral states. This model is inspired by molecular dynamics modeling that defines the foraging reorientation rate as a decaying parameter. By stochastically sampling from the time interval probability distribution defined by this rate, both abrupt and gradual changes to reorientation rates can occur, matching experimentally observed results. Crucially, this model does not depend on behavioral states or information accumulation. Even though abrupt behavioral changes do occur, they are not necessarily indicative of abrupt changes in behavioral states, especially when abrupt changes are not universally observed in the population.
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spelling doaj-art-afb5854925dc44d38ff4641b3a2f62862025-08-20T02:46:24ZengeLife Sciences Publications LtdeLife2050-084X2025-07-011410.7554/eLife.104972A stochastic explanation for observed local-to-global foraging states in Caenorhabditis elegansAndrew Margolis0https://orcid.org/0009-0006-9817-7147Andrew Gordus1https://orcid.org/0000-0002-5550-0286Department of Biology, Johns Hopkins University, Baltimore, United States; David Geffen School of Medicine, University of California, Los Angeles, United StatesDepartment of Biology, Johns Hopkins University, Baltimore, United States; Solomon H. Snyder Department of Neuroscience, Johns Hopkins University, Baltimore, United StatesAbrupt changes in behavior can often be associated with changes in underlying behavioral states. When placed off food, the foraging behavior of C. elegans can be described as a change between an initial local-search behavior characterized by a high rate of reorientations, followed by a global-search behavior characterized by sparse reorientations. This is commonly observed in individual worms, but when numerous worms are characterized, only about half appear to exhibit this behavior. We propose an alternative model that predicts both abrupt and continuous changes to reorientation that do not rely on behavioral states. This model is inspired by molecular dynamics modeling that defines the foraging reorientation rate as a decaying parameter. By stochastically sampling from the time interval probability distribution defined by this rate, both abrupt and gradual changes to reorientation rates can occur, matching experimentally observed results. Crucially, this model does not depend on behavioral states or information accumulation. Even though abrupt behavioral changes do occur, they are not necessarily indicative of abrupt changes in behavioral states, especially when abrupt changes are not universally observed in the population.https://elifesciences.org/articles/104972behaviorinternal statestochasticGillespieforaging
spellingShingle Andrew Margolis
Andrew Gordus
A stochastic explanation for observed local-to-global foraging states in Caenorhabditis elegans
eLife
behavior
internal state
stochastic
Gillespie
foraging
title A stochastic explanation for observed local-to-global foraging states in Caenorhabditis elegans
title_full A stochastic explanation for observed local-to-global foraging states in Caenorhabditis elegans
title_fullStr A stochastic explanation for observed local-to-global foraging states in Caenorhabditis elegans
title_full_unstemmed A stochastic explanation for observed local-to-global foraging states in Caenorhabditis elegans
title_short A stochastic explanation for observed local-to-global foraging states in Caenorhabditis elegans
title_sort stochastic explanation for observed local to global foraging states in caenorhabditis elegans
topic behavior
internal state
stochastic
Gillespie
foraging
url https://elifesciences.org/articles/104972
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AT andrewmargolis stochasticexplanationforobservedlocaltoglobalforagingstatesincaenorhabditiselegans
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